Argynnini

Niklas Wahlberg

Based on Markku Savela's pages and Lamas, G. 2004: Atlas of Neotropical Lepidoptera. Checklist: Part 4A, Hesperioidea - Papilionoidea. Scientific Publishers, Gainsville, Florida.

TRIB Argynnini Swainson, 1833

GENU Euptoieta Doubleday, 1848

SPEC Euptoieta bogotana Staudinger, 1885

SSYN Euptoieta colombia Strecker, 1899

SPEC Euptoieta claudia (Cramer, 1775)

SUBS Euptoieta claudia claudia (Cramer, 1775)

ORIG Papilio claudia Cramer, 1775
SSYN Papilio thais Doubleday, 1848

SUBS Euptoieta claudia daunius (Herbst, 1798)

ORIG Papilio daunius Herbst, 1798
SSYN Euptoieta claudia f. dodgei Gunder, 1927
SSYN Euptoieta claudia f. fumosa Field, 1936
SSYN Euptoieta claudia f. albaclaudia Field, 1936

SPEC Euptoieta hegesia (Cramer, 1779)

SUBS Euptoieta hegesia hegesia (Cramer, 1779)

ORIG Papilio hegesia Cramer, 1779
SSYN Euptoieta hegesia karibica Stichel, 1938
SSYN Euptoieta hegesia domingensis Cucurullo, 1959

SUBS Euptoieta hegesia meridiana Stichel, 1938

SSYN Euptoieta hegesia hoffmanni Comstock, 1944

SPEC Euptoieta hortensia (Blanchard, 1852)

ORIG Argynnis hortensia Blanchard, 1852
SSYN Argynnis valdiviana Philippi, 1859
SSYN Euptoieta ramirezi Giacomelli, 1926
SSYN Euptoieta claudia ab. infuscata Hayward, 1940

SPEC Euptoieta perdistincta Hall, 1930

SSYN Brenthis hana Clark, 1936

SPEC Euptoieta poasina Schaus, 1913
SPEC Euptoieta sunides (Hewitson, 1877)

ORIG Argynnis sunides Hewitson, 1877

SPEC Euptoieta thekla Hall, 1919
SPEC Euptoieta n. sp. Lamas, MS

GENU Yramea Reuss, 1920

GSYN Chilargynnis Bryk, 1944

SPEC Yramea cytheris (Drury, 1773)

SUBS Yramea cyhteris cytheris (Drury, 1773)

ORIG Papilio cytheris Drury, 1773
SSYN Argynnis cytheris falklandica Watkins, 1924

SUBS Yramea cyhteris siga (Geyer, 1832)

ORIG Argynnis siga Geyer, 1832
SSYN Argynnis anna Blanchard, 1852
SSYN Argynnis chilensis Reed, 1877
SSYN Chilargynnis cytheris subtusviola Bryk, 1944

SPEC Yramea inca (Staudinger, 1894)

SUBS Yramea inca inca (Staudinger, 1894)

ORIG Argynnis inca Staudinger, 1894

SUBS Yramea inca cora (Lehmann, 1913)

ORIG Argynnis cora Lehmann, 1913

SPEC Yramea lathonioides (Blanchard, 1852)

ORIG Argynnis lathonioides Blanchard, 1852
SSYN Argynnis dexamene Boisduval, 1859
SSYN Argynnis dioides Boisduval, 1859
SSYN Argynnis montana Reed, 1877
SSYN Argynnis euterpe Reed, 1877
SSYN Argynnis darwini Staudinger, 1899

SPEC Yramea modesta (Blanchard, 1852)

ORIG Argynnis modesta Blanchard, 1852

SPEC Yramea sobrina (Weymer, 1890)

ORIG Argynnis sobrina Weymer, 1890
SSYN Boloria tessellata Rosenberg & Talbot, 1914
SSYN Yramea blancasi Hayward, 1960

SPEC Yramea lynx Lamas & Grados, 2004

GENU Argynnis Fabricius, 1807

GSYN Pandoriana Warren, 1942
GSYN Childrena Hemming, 1943
GSYN Eudryas Reuss, 1926 (preocc. Eudryas Boisduval, 1836; Eudryas Harris, 1841; Eudryas Fitzinger, 1843; Eudryas Gistl, 1848)
GSYN Argyreus Scopoli, 1777
GSYN Acidalia Hübner, 1819
GSYN Argyrea Billberg, 1820
GSYN Mimargyra Reuss, 1922
GSYN Damora Nordmann, 1851
GSYN Argyronome Hübner, 1819

SPEC Argynnis paphia (Linnaeus, 1758)

ORIG Papilio paphia Linnaeus, 1758

SUBS Argynnis paphia paphia (Linnaeus, 1758)
SUBS Argynnis paphia valesina Esper
SUBS Argynnis paphia dives (Oberthür, 1908)
SUBS Argynnis paphia formosicola Matsumura
SUBS Argynnis paphia neopaphia Fruhstorfer
SUBS Argynnis paphia megalegoria Fruhstorfer, 1907

SPEC Argynnis sagana Doubleday, 1847

SUBS Damora sagana sagana (Doubleday, 1847)
SUBS Damora sagana liane (Fruhstorfer, 1907)
SUBS Damora sagana relicta Korshunov, 1984
SUBS Damora sagana paulina (Nordman, 1851)

SPEC Argynnis anadyomene Felder & Felder, 1862
SPEC Argynnis pandora (Denis & Schiffermüller, 1775)

ORIG Papilio pandora Denis & Schiffermüller, 1775
SSYN Papilio maja Cramer, 1775
SSYN Pandoriana pandora (Denis & Schiffermüller, 1775)

SUBS Argynnis pandora pandora (Denis & Schiffermüller, 1775)
SUBS Argynnis pandora seitzi Fruhstorfer, 1908

SPEC Argynnis childreni Gray, 1831

SUBS Argynnis childreni childreni Gray, 1831
SUBS Argynnis childreni sakontala Kollar
SUBS Argynnis childreni caesarea Fruhstorfer

SPEC Argynnis zenobia (Leech, 1890)

SUBS Argynnis zenobia zenobia (Leech, 1890)
SUBS Argynnis zenobia penelope (Staudinger)

SPEC Argynnis hyperbius (Linnaeus, 1763)

ORIG Papilio hyperbius Linnaeus, 1763
SSYN Papilio niphe Linnaeus, 1767 (unnec. repl. for Papilio hyperbius Linnaeus, 1763)

SUBS Argynnis hyperbius hyperbius (Linnaeus, 1763)
SUBS Argynnis hyperbius taprobana Moore
SUBS Argynnis hyperbius castetsi Oberthür
SUBS Argynnis hyperbius sumatrensis (Fruhstorfer, 1905)
SUBS Argynnis hyperbius javanica Oberthür
SUBS Argynnis hyperbius sagada Fruhstorfer
SUBS Argynnis hyperbius inconstans (Butler, 1873)
SUBS Argynnis hyperbius neumanni Rothschild & Jordan

SPEC Argynnis laodice (Pallas, 1771)

ORIG Papilio laodice Pallas, 1771

SUBS Argynnis laodice laodice (Pallas, 1771)
SUBS Argynnis laodice rudra (Moore, 1857)
SUBS Argynnis laodice samana Fruhstorfer, 1902
SUBS Argynnis laodice japonica (Ménétriés)
SUBS Argynnis laodice huochengice Huang & Murayama

SPEC Argynnis kuniga Chou & Tong, 1994
SPEC Argynnis ruslana Motschulsky, 1866

GENU Speyeria Scudder, 1872

GSYN Semnopsyche Scudder, 1875
GSYN Mesoacidalia Reuss, 1926
GSYN Proacidalia Reuss, 1926

SPEC Speyeria aglaja (Linnaeus, 1758)

ORIG Papilio aglaja Linnaeus, 1758
SSYN Papilio pasilhoe Linnaeus, 1767
SSYN Papilio charlotta Haworth, 1802
SSYN Mesoacidalia aglaja (Linnaeus, 1758)

SUBS Speyeria aglaja aglaja (Linnaeus, 1758)
SUBS Speyeria aglaja lyauteyi (Oberthür, 1920)
SUBS Speyeria aglaja excelsior (Rothschild, 1933)
SUBS Speyeria aglaja vitatha (Moore)
SUBS Speyeria aglaja clavimacula (Matsumura)
SUBS Speyeria aglaja bessa (Fruhstorfer, 1907)

SPEC Speyeria clara (Blanchard, 1844)

ORIG Argynnis clara Blanchard, 1844

SPEC Speyeria alexandra Ménétriés, 1855
SPEC Speyeria claudia Fawcett
SPEC Speyeria diana (Cramer, 1777)

ORIG Papilio diana Cramer, 1777

SPEC Speyeria cybele (Fabricius, 1775)

ORIG Papilio cybele (Fabricius, 1775)

SUBS Speyeria cybele cybele (Fabricius, 1775)
SUBS Speyeria cybele novascotiae (McDunnough, 1935)
SUBS Speyeria cybele krautwurmi (Holland, 1931)
SUBS Speyeria cybele pseudocarpenteri (F. & R. Chermock, 1940)
SUBS Speyeria cybele carpenterii (Edwards, 1876)
SUBS Speyeria cybele charlottii (Barnes, 1897)
SUBS Speyeria cybele letona dos Passos & Grey, 1945
SUBS Speyeria cybele pugetensis F. Chermock & Frechin, 1947
SUBS Speyeria cybele leto (Behr, 1862)

SPEC Speyeria aphrodite (Fabricius, 1787)

ORIG Papilio aphrodite (Fabricius, 1787)

SUBS Speyeria aphrodite aphrodite (Fabricius, 1787)
SUBS Speyeria aphrodite winni (Gunder, 1932)
SUBS Speyeria aphrodite alcestis (Edwards, 1877)
SUBS Speyeria aphrodite manitoba (F. & R. Chermock, 1940)
SUBS Speyeria aphrodite whitehousei (Gunder, 1932)
SUBS Speyeria aphrodite columbia (Hy. Edwards, 1877)
SUBS Speyeria aphrodite ethene (Hemming, 1933)
SUBS Speyeria aphrodite byblis (Barnes & Benjamin, 1926)

SPEC Speyeria idalia (Drury, 1773)

ORIG Papilio idalia Drury, 1773

SPEC Speyeria nokomis (Edwards, 1862)

SUBS Speyeria nokomis nokomis (Edwards, 1862)
SUBS Speyeria nokomis nitocris (Edwards, 1874)
SUBS Speyeria nokomis coerulescens (Holland, 1900)
SUBS Speyeria nokomis apacheana (Skinner, 1918)

SPEC Speyeria edwardsii (Reakirt, 1866)
SPEC Speyeria coronis (Behr, 1864)

SUBS Speyeria coronis coronis (Behr, 1864)
SUBS Speyeria coronis hennei (Gunder, 1934)
SUBS Speyeria coronis semiramis (Edwards, 1886)
SUBS Speyeria coronis gunderi (Comstock, 1925)
SUBS Speyeria coronis simaetha dos Passos & Grey, 1945
SUBS Speyeria coronis snyderi (Skinner, 1897)
SUBS Speyeria coronis halcyone (Edwards, 1869)

SPEC Speyeria zerene (Boisduval, 1852)

SUBS Speyeria zerene zerene (Boisduval, 1852)
SUBS Speyeria zerene conchyliata (Comstock, 1925)
SUBS Speyeria zerene gloriosa Moeck, 1957
SUBS Speyeria zerene sordida (W. G. Wright, 1905)
SUBS Speyeria zerene malcomi (Comstock, 1920)
SUBS Speyeria zerene carolae (dos Passos & Grey, 1942)
SUBS Speyeria zerene hippolyta (Edwards, 1879)
SUBS Speyeria zerene behrensii (Edwards, 1869)
SUBS Speyeria zerene myrtleae dos Passos & Grey, 1945
SUBS Speyeria zerene bremnerii (Edwards, 1872)
SUBS Speyeria zerene picta (McDunnough, 1924)
SUBS Speyeria zerene garretti (Gunder, 1932)
SUBS Speyeria zerene sinope dos Passos & Grey, 1945
SUBS Speyeria zerene platina (Skinner, 1897)
SUBS Speyeria zerene pfoutsi (Gunder, 1933)
SUBS Speyeria zerene cynna dos Passos & Grey, 1945

SPEC Speyeria callippe (Boisduval, 1852)

SUBS Speyeria callippe callippe (Boisduval, 1852)
SUBS Speyeria callippe comstocki (Gunder, 1925)
SUBS Speyeria callippe liliana (Hy. Edwards, 1877)
SUBS Speyeria callippe semivirida (McDunnough, 1924)
SUBS Speyeria callippe elaine dos Passos & Grey, 1945
SUBS Speyeria callippe rupestris (Behr, 1863)
SUBS Speyeria callippe juba (Boisduval, 1869)
SUBS Speyeria callippe laura (Edwards, 1879)
SUBS Speyeria callippe sierra dos Passos & Grey, 1945
SUBS Speyeria callippe nevadensis (Edwards, 1870)
SUBS Speyeria callippe macaria (Edwards, 1877)
SUBS Speyeria callippe laurina (W. G. Wright, 1905)
SUBS Speyeria callippe harmonia dos Passos & Grey, 1945
SUBS Speyeria callippe meadii (Edwards, 1872)
SUBS Speyeria callippe gallatini (McDunnough, 1929)
SUBS Speyeria callippe calgariana (McDunnough, 1924)

SPEC Speyeria egleis (Behr, 1863)

SUBS Speyeria egleis egleis (Behr, 1863)
SUBS Speyeria egleis clemencei (Comstock, 1925)
SUBS Speyeria egleis atossa (Edwards, 1890)
SUBS Speyeria egleis tehachapina (Comstock, 1920)
SUBS Speyeria egleis oweni (Edwards, 1892)
SUBS Speyeria egleis linda (dos Passos & Grey, 1942)
SUBS Speyeria egleis macdunnoughi (Gunder, 1932)
SUBS Speyeria egleis albrighti (Gunder, 1932)
SUBS Speyeria egleis utahensis (Skinner, 1919)
SUBS Speyeria egleis toiyabe Howe, 1975
SUBS Speyeria egleis secreta dos Passos & Grey, 1945

SPEC Speyeria adiaste (Edwards, 1864)
SPEC Speyeria atlantis (Edwards, 1862)

SUBS Speyeria atlantis atlantis (Edwards, 1862)
SUBS Speyeria atlantis canadensis (dos Passos, 1935)
SUBS Speyeria atlantis hollandi (F. & R. Chermock, 1940)
SUBS Speyeria atlantis hesperis (Edwards, 1864)
SUBS Speyeria atlantis nikias (Ehrmann, 1917)
SUBS Speyeria atlantis dorothea Moeck, 1947
SUBS Speyeria atlantis nausicaa (Edwards, 1874)
SUBS Speyeria atlantis schellbachi Garth, 1949
SUBS Speyeria atlantis chitone (Edwards, 1879)
SUBS Speyeria atlantis wasatchia dos Passos & Grey, 1945
SUBS Speyeria atlantis greyi (Moeck, 1950)
SUBS Speyeria atlantis tetonia dos Passos & Grey, 1945
SUBS Speyeria atlantis viola dos Passos & Grey, 1945
SUBS Speyeria atlantis dodgei (Gunder, 1931)
SUBS Speyeria atlantis irene (Boisduval, 1869)
SUBS Speyeria atlantis electa (Edwards, 1878)
SUBS Speyeria atlantis lurana dos Passos & Grey, 1945
SUBS Speyeria atlantis hutchinsi dos Passos & Grey, 1947
SUBS Speyeria atlantis beani (Barnes & Benjamin, 1926)
SUBS Speyeria atlantis lais (Edwards, 1884)
SUBS Speyeria atlantis dennisi dos Passos & Grey, 1947

SPEC Speyeria hydaspe (Boisduval, 1869)

SUBS Speyeria hydaspe hydaspe (Boisduval, 1869)
SUBS Speyeria hydaspe virdicornis (Comstock, 1925)
SUBS Speyeria hydaspe purpurascens (Hy. Edwards, 1877)
SUBS Speyeria hydaspe minor dos Passos & Grey, 1947
SUBS Speyeria hydaspe rhodope (Edwards, 1874)
SUBS Speyeria hydaspe sakuntala (Skinner, 1911)
SUBS Speyeria hydaspe conquista dos Passos & Grey, 1945

SPEC Speyeria mormonia (Boisduval, 1869)

SUBS Speyeria mormonia mormonia (Boisduval, 1869)
SUBS Speyeria mormonia bischoffii (Edwards, 1870)
SUBS Speyeria mormonia opis (Edwards, 1874)
SUBS Speyeria mormonia jesmondensis dos Passos & Grey, 1947
SUBS Speyeria mormonia washingtonia (Barnes & McDunnough, 1913)
SUBS Speyeria mormonia erinna (Edwards, 1883)
SUBS Speyeria mormonia arge (Strecker, 1878)
SUBS Speyeria mormonia artonis (Gunder, 1932)
SUBS Speyeria mormonia eurynome (Edwards, 1872)
SUBS Speyeria mormonia luski (Barnes & McDunnough, 1913)

GENU Fabriciana Reuss, 1920

GSYN Profabriciana Reuss, 1926
GSYN Protodryas Reuss, 1928

SPEC Fabriciana niobe (Linnaeus, 1758)

ORIG Papilio niobe Linnaeus, 1758

SUBS Fabriciana niobe niobe (Linnaeus, 1758)
SUBS Fabriciana niobe changaica Reuss, 1922
SUBS Fabriciana niobe ornata (Staudinger)
SUBS Fabriciana niobe orientalis (Alphéraky, 1881)

SPEC Fabriciana coredippe Leech, 1892
SPEC Fabriciana auresiana (Fruhstorfer, 1908)

ORIG Argynnis adippe auresiana Fruhstorfer, 1908

SUBS Fabriciana auresiana auresiana (Fruhstorfer, 1908)
SUBS Fabriciana auresiana hassani Weiss, 1979
SUBS Fabriciana auresiana astrifera Higgins, 1965

SPEC Fabriciana adippe (Denis & Schiffermüller, 1775)

ORIG Papilio adippe Denis & Schiffermüller, 1775
SSYN Papilio cydippe Linnaeus, 1761
SSYN Argynnis adippe (Denis & Schiffermüller, 1775)

SUBS Fabriciana adippe adippe (Denis & Schiffermüller, 1775)
SUBS Fabriciana adippe ornatissima (Leech, 1892)
SUBS Fabriciana adippe vorax (Butler, 1871)
SUBS Fabriciana adippe tianshanica (Staudinger)
SUBS Fabriciana adippe zarevna Fruhstorfer, 1912
SUBS Fabriciana adippe xanthodippe (Fixsen)
SUBS Fabriciana adippe taurica (Staudinger)
SUBS Fabriciana adippe taliana Reuss, 1922
SUBS Fabriciana adippe pallescens (Butler, 1873)
SUBS Fabriciana adippe chayuensis Huang, 2001

SPEC Fabriciana xipe (Groum-Grshimailo, 1891)

SUBS Fabriciana xipe xipe (Groum-Grshimailo, 1891)
SUBS Fabriciana xipe gyala (Tytler, 1940)

SPEC Fabriciana elisa (Godart, 1823)

ORIG Argynnis elisa Godart, 1823

SPEC Fabriciana argyrospilata Kotsch, 1938
SPEC Fabriciana nerippe (Felder, 1862)

SUBS Fabriciana nerippe nerippe (Felder, 1862)
SUBS Fabriciana nerippe nerippina (Fruhstorfer)

SPEC Fabriciana kamala (Moore, 1857)

ORIG Argynnis kamala Moore, 1857

SPEC Fabriciana jainadeva (Moore, 1864)

ORIG Argynnis jainadeva Moore, 1864
SSYN Profabriciana jainadeva (Moore, 1864)

SPEC Fabriciana funebris Oberthür

GENU Brenthis Hübner, 1819

SPEC Brenthis ino (Rottemburg, 1775)

ORIG Papilio ino (Rottemburg, 1775)

SUBS Brenthis ino ino (Rottemburg, 1775)
SUBS Brenthis ino paidicus Fruhstorfer, 1907
SUBS Brenthis ino acrita Fruhstorfer
SUBS Brenthis ino amurensis (Staudinger)
SUBS Brenthis ino discolus (Fruhstorfer)

SPEC Brenthis hecate (Denis & Schiffermüller, 1775)

ORIG Papilio hecate Denis & Schiffermüller, 1775

SUBS Brenthis hecate hecate (Denis & Schiffermüller, 1775)
SUBS Brenthis hecate alaica Staudinger
SUBS Brenthis hecate warren Kudrna, 1974

SPEC Brenthis daphne (Denis & Schiffermüller, 1775)

ORIG Papilio daphne Denis & Schiffermüller, 1775

SUBS Brenthis daphne daphne (Denis & Schiffermüller, 1775)
SUBS Brenthis daphne ochroleuca (Fruhstorfer)

GENU Issoria Hübner, 1819

GSYN Rathora Moore, 1900
GSYN Prokuekenthaliella Reuss, 1927
GSYN Kuekenthaliella Reuss, 1921

SPEC Issoria lathonia (Linnaeus, 1758)

ORIG Papilio lathonia Linnaeus, 1758

SUBS Issoria lathonia lathonia (Linnaeus, 1758)
SUBS Issoria lathonia issaea (Gray, 1857)

SPEC Issoria hanningtoni Elwes, 1889

SUBS Issoria hanningtoni hanningtoni Elwes, 1889
SUBS Issoria hanningtoni jeanneli Bernardi, 1968
SUBS Issoria hanningtoni imatonga Riley

SPEC Issoria smaragdifera (Butler, 1895)

ORIG Argynnis smaragdifera Butler, 1895

SUBS Issoria smaragdifera smaragdifera (Butler, 1895)
SUBS Issoria smaragdifera reducta Carcasson

SPEC Issoria baumanni Rebel, 1894

SUBS Issoria baumanni baumanni Rebel, 1894
SUBS Issoria baumanni excelsior (Butler, 1896)
SUBS Issoria baumanni katangae Neave

SPEC Issoria uganda Rob.
SPEC Issoria sinha (Kollar)
SPEC Issoria gemmata (Butler, 1881)

ORIG Argynnis gemmata Butler, 1881

SUBS Issoria gemmata gemmata (Butler, 1881)
SUBS Issoria gemmata wui Huang, 1998

SPEC Issoria baileyi Huang, 1998
SPEC Issoria eugenia (Eversmann, 1847)

SUBS Issoria eugenia eugenia (Eversmann, 1847)
SUBS Issoria eugenia montana (Bang-Haas, 1906)
SUBS Issoria eugenia rhea (Groum-Grshimailo, 1891)
SUBS Issoria eugenia genia (Fruhstorfer, 1903)
SUBS Issoria eugenia pulchella Huang, 2001
SUBS Issoria eugenia tibetana Huang, 1998

SPEC Issoria altissima (Elwes, 1882)
SPEC Issoria mackinnonii (de Nicéville, 1891)

GENU Boloria Moore, 1900

SPEC Boloria pales (Denis & Schiffermüller, 1775)

ORIG Papilio pales Denis & Schiffermüller, 1775

SUBS Boloria pales pales (Denis & Schiffermüller, 1775)
SUBS Boloria pales palina (Fruhstorfer, 1903)
SUBS Boloria pales eupales (Fruhstorfer, 1903)
SUBS Boloria pales shambhala Kocman, 1999
SUBS Boloria pales nirvana Kocman, 1999
SUBS Boloria pales sifanica (Grum-Grshimailo, 1891)

SPEC ?Boloria generator Staudinger, 1886
SPEC Boloria napaea (Hoffmannsegg, 1804)

SUBS Boloria napaea napaea (Hoffmannsegg, 1804)
SUBS Boloria napaea altaica (Groum-Grshimailo, 1893)
SUBS Boloria napaea alaskensis (Holland, 1900)
SUBS Boloria napaea nearctica Verity, 1932
SUBS Boloria napaea halli Klots, 1940

SPEC Boloria frigidalis Warren, 1944
SPEC Boloria aquilonaris (Stichel, 1980)

SUBS Boloria aquilonaris aquilonaris (Stichel, 1980)
SUBS Boloria aquilonaris banghaasi (Seitz, 1909)

SPEC Boloria graeca (Staudinger, 1870)

GENU Proclossiana Reuss, 1926

SPEC Proclossiana eunomia (Esper, 1799)

ORIG Papilio eunomia Esper, 1799
SSYN Papilio aphirape Hübner, 1799
SSYN Boloria eunomia (Esper, 1799)
SSYN Clossiana eunomia (Esper, 1799)

SUBS Proclossiana eunomia eunomia (Esper, 1799)
SUBS Proclossiana eunomia ossiana (Herbst, 1800)
SUBS Proclossiana eunomia triclaris (Hübner, 1821)
SUBS Proclossiana eunomia caelestis (Hemming, 1933)
SUBS Proclossiana eunomia dawsoni (Barnes & McDunnough, 1916)
SUBS Proclossiana eunomia nichollae (Barnes & Benjamin, 1926)
SUBS Proclossiana eunomia laddi (Klots, 1940)
SUBS Proclossiana eunomia ursadentis (Ferris & Groothuis, 1971)
SUBS Proclossiana eunomia denali (Klots, 1940)
SUBS Proclossiana eunomia acidalia Boeber, 1809

GENU Clossiana Reuss, 1920

SPEC Clossiana selene (Denis & Schiffermüller, 1775)

ORIG Papilio selene Denis & Schiffermüller, 1775
SSYN Boloria selene

SUBS Clossiana selene selene (Denis & Schiffermüller, 1775)
SUBS Clossiana selene thalia Hübner, 1790
SUBS Clossiana selene dilutior Fixsen, 1887
SUBS Clossiana selene chibiana Matsumura, 1927
SUBS Clossiana selene perryi Butler, 1882
SUBS Clossiana selene myrina (Cramer, 1777)
SUBS Clossiana selene nebraskensis (Holland, 1928)
SUBS Clossiana selene sabulicollis (Kohler, 1977)
SUBS Clossiana selene tolladensis (Barnes & Benjamin, 1925)
SUBS Clossiana selene albequina (Holland, 1928)
SUBS Clossiana selene atrocostalis (Huard, 1927)
SUBS Clossiana selene terraenovae (Holland, 1928)

SPEC Clossiana erubescens (Staudinger, 1901)
SPEC Clossiana selenis (Eversmann, 1837)

SUBS Clossiana selenis selenis (Eversmann, 1837)
SUBS Clossiana selenis sibirica Ershov, 1870

SPEC Clossiana angarensis (Erschoff, 1870)

SUBS Clossiana angarensis angarensis (Erschoff, 1870)
SUBS Clossiana angarensis samkoi Sheljuzhko, 1931
SUBS Clossiana angarensis alticoa Sushkin & Tshetverikov, 107
SUBS Clossiana angarensis herzi Wnukowsky, 1927
SUBS Clossiana angarensis miakei Matsumura, 1919

SPEC Clossiana oscarus (Eversmann, 1844)

SUBS Clossiana oscarus oscarus (Eversmann, 1844)
SUBS Clossiana oscarus australis Graeser, 1888

SPEC Clossiana freija (Thunberg, 1791)

ORIG Papilio freija Thunberg, 1791

SUBS Clossiana freija freija (Thunberg, 1791)
SUBS Clossiana freija jakutensis Wnukowsky, 1927
SUBS Clossiana freija pallida Elwes, 1899
SUBS Clossiana freija tarquinius (Curtis, 1835)
SUBS Clossiana freija natazhati (Gibson, 1920)
SUBS Clossiana freija nabokovi (D. Stallings & Turner, 1947)
SUBS Clossiana freija browni (Higgins, 1953)
SUBS Clossiana freija pallida Elwes, 1899

SPEC Clossiana polaris (Boisduval, 1828)

SUBS Clossiana polaris polaris (Boisduval, 1828)
SUBS Clossiana polaris stellata (Masters, 1972)

SPEC Clossiana thore (Hübner, 1803)

SUBS Clossiana thore thore (Hübner, 1803)
SUBS Clossiana thore borealis (Staudinger, 1861)
SUBS Clossiana thore transuralensis Sheljuzhko, 1931
SUBS Clossiana thore herculea Fruhstorfer, 1907
SUBS Clossiana thore hyperusia Fruhstorfer, 1907
SUBS Clossiana thore meridionalis Kardakov, 1928

SPEC Clossiana hakutozana (Matsumura, 1927)
SPEC Clossiana natazhati Gibson, 1920
SPEC Clossiana frigga (Thunberg, 1791)

ORIG Papilio frigga Thunberg, 1791

SUBS Clossiana frigga frigga (Thunberg, 1791)
SUBS Clossiana frigga alpestris Elwes, 1899
SUBS Clossiana frigga maritima Kardakov, 1928
SUBS Clossiana frigga saga (Staudinger, 1861)
SUBS Clossiana frigga alaskensis (Lehmann, 1913)
SUBS Clossiana frigga sagata (Barnes & Benjamin, 1923)

SPEC Clossiana dia (Linnaeus, 1767)

ORIG Papilio dia Linnaeus, 1767

SUBS Clossiana dia dia (Linnaeus, 1767)
SUBS Clossiana dia alpina Elwes, 1899
SUBS Clossiana dia setania Fruhstorfer, 1909

SPEC Clossiana gong (Oberthür, 1884)

SUBS Clossiana gong gong (Oberthür, 1884)
SUBS Clossiana gong splendens Murayama
SUBS Clossiana gong xizangensis Huang, 2000

SPEC Clossiana jerdoni (Lang, 1868)

ORIG Argynnis jerdoni Lang, 1868

SPEC Clossiana improba (Butler, 1877)

SUBS Clossiana improba improba (Butler, 1877)
SUBS Clossiana improba improbula (Bryk, 1920)
SUBS Clossiana improba youngi (Holland, 1900)
SUBS Clossiana improba acrocnema

SPEC Clossiana titania (Esper, 1793)

ORIG Papilio titania Esper, 1793
SSYN Papilio amathusia Esper, 1784

SUBS Clossiana titania titania (Esper, 1793)
SUBS Clossiana titania bivina Fruhstorfer, 1909
SUBS Clossiana titania staudingeri Wnukowsky, 1929
SUBS Clossiana titania miyakei Matsumura, 1919
SUBS Clossiana titania boisduvalii (Duponchel, 1832)
SUBS Clossiana titania rainieri (Barnes & McDunnough, 1913)
SUBS Clossiana titania grandis (Barnes & McDunnough, 1916)
SUBS Clossiana titania montina (Scudder, 1863)
SUBS Clossiana titania ingens (Barnes & McDunnough, 1918)
SUBS Clossiana titania helena (Edwards, 1871)

SPEC Clossiana euphrosyne (Linnaeus, 1758)

ORIG Papilio euphrosyne Linnaeus, 1758

SUBS Clossiana euphrosyne euphrosyne (Linnaeus, 1758)
SUBS Clossiana euphrosyne nephele Herrich-Schaeffer, 1846
SUBS Clossiana euphrosyne oprhanus Fruhstorfer, 1907
SUBS Clossiana euphrosyne kamtschadalus Seitz, 1908

SPEC Clossiana chariclea (Schneider, 1794)

SUBS Clossiana chariclea chariclea (Schneider, 1794)
SUBS Clossiana chariclea arctica (Zetterstedt, 1839)
SUBS Clossiana chariclea butleri (Edwards, 1883)

SPEC Clossiana bellona (Fabricius, 1775)

ORIG Papilio bellona Fabricius, 1775

SUBS Clossiana bellona bellona (Fabricius, 1775)
SUBS Clossiana bellona toddi (Holland, 1928)
SUBS Clossiana bellona jenistae (D. Stallings & Turner, 1947)

SPEC Clossiana kriemhild (Strecker, 1879)
SPEC Clossiana epithore (Edwards, 1864)

SUBS Clossiana epithore epithore (Edwards, 1864)
SUBS Clossiana epithore chermocki (E. & S. Perkins, 1966)
SUBS Clossiana epithore borealis (E. Perkins, 1973)
SUBS Clossiana epithore sierra (E. Perkins, 1973)

SPEC Clossiana alberta (Edwards, 1890)

SUBS Clossiana alberta alberta (Edwards, 1890)
SUBS Clossiana alberta kurentzovi Wyatt, 1961

SPEC Clossiana astarte (Doubleday & Hewitson, 1847)
SPEC Clossiana tritonia (de Boeber, 1812)
SPEC Clossiana distincta (Gibson, 1920)

SUBS Clossiana distincta distincta (Gibson, 1920)
SUBS Clossiana distincta machati Korshunov, 1987
SUBS Clossiana distincta dubatolovi Korshunov, 1987
SUBS Clossiana distincta tschukotkensis Wyatt, 1961

SPEC Clossiana matveevi Gorbunov & Korshunov, 1995
SPEC Clossiana erda (Christoph, 1892)

SUBS Clossiana erda erda (Christoph, 1892)
SUBS Clossiana erda dulkeiti Kurenzov, 1970

SPEC Clossiana elatus (Staudinger, 1892)
SPEC Clossiana iphigenia (Graeser, 1888)

SUBS Clossiana iphigenia iphigenia (Graeser, 1888)
SUBS Clossiana iphigenia sachalinensis Matsumura, 1925

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