Melitaeini

Niklas Wahlberg

Euphydryas based on Higgins, L. G. 1950: A descriptive catalogue of the Palearctic Euphydryas (Lepidoptera: Rhopalocera). - Trans. R. Ent. Soc. Lond. 101: 435-487.

Melitaea based on Higgins, L. G. 1941: An illustrated catalogue of the Palearctic Melitaea (Lep. Rhopalocera). - Trans. R. Ent. Soc. Lond. 91: 175-365. and
Higgins, L. G. 1955: A descriptive catalogue of the genus Mellicta Billberg (Lepidoptera: Nymphalidae) and its species, with supplementary notes on the genera Melitaea and Euphydryas. - Trans. R. Ent. Soc. Lond. 10: 61-127.
Achtelik, G. 1999: Vier neue arten aus der "Melitaea sultanensis-gruppe" (Staudinger, 1886) (Lepidoptera, Nymphalidae). - Atalanta 30: 63-86.
Kolesnichenko, K. A. 1999: A review of the East-Palaearctic taxa of the Melitaea didyma (Esper, 1779)-group 1. The M. ala Staudinger, 1881-M. chitralensis Moore, 1901-complex (Lepidoptera, Nymphalidae). - Atalanta 30: 87-117.

Chlosynina based on Kons, H. L. Jr. 2000: Phylogenetic studies of the Melitaeini (Lepidoptera: Nymphalidae: Nymphalinae) and a revision of the genus Chlosyne Butler. - PhD thesis, University of Florida, Gainesville, 799 pp. Available from UMI.

Phyciodina based on Higgins, L. G. 1981. A revision of Phyciodes Hübner and related genera, with a review of the classification of the Melitaeinae (Lepidoptera: Nymphalidae). Bull. Brit. Mus. Nat. Hist. 43: 77-243. and
Scott, J. A. 1994. Biology and systematics of Phyciodes (Phyciodes). Papilio (New Series) 7: 1-120. and
Scott, J. A. 1998. Phyciodes (Phyciodes): new discoveries, new subspecies, and convergence. - Papilio (New Series) 10: 1-42.

SUBT Euphydryina Higgins, 1978
GENU Euphydryas Scudder, 1872

GSYN Occidryas Higgins, 1978
GSYN Eurodryas Higgins, 1978
GSYN HypodryasHiggins, 1978

SPEC Euphydryas aurinia (Rottemburg, 1775)

ORIG Papilio aurinia Rottemburg, 1775
SSYN Papilio artemis Schiffermüller, 1775
SSYN Papilio lye Bergstrasser, 1780
SSYN Papilio koloswarensis Piller & Mitterpacher, 1783
SSYN Papilio matutina Thunberg, 1791
SSYN Eurodryas aurinia (Rottemburg, 1775)

SUBS Euphydryas aurinia aurinia (Rottemburg, 1775)
SUBS Euphydryas aurinia beckeri (Lederer, 1853)

ORIG Melitaea beckeri Lederer, 1853
SSSP Melitaea aurinia f. iberica Oberthür, 1881
SSSP Melitaea aurinia ellisoni Rungs, 1950
SSSP Melitaea aurinia kandarensis Rungs, 1950

SUBS Euphydryas aurinia barraguei (Betz, 1956)

ORIG Melitaea aurinia barraguei Betz, 1956

SUBS Euphydryas aurinia sareptensis (Staudinger, 1879)

ORIG Melitaea aurinia sareptensis Staudinger, 1879

SUBS Euphydryas aurinia banghaasi (Seitz, 1907)
SUBS Euphydryas aurinia asiatica (Staudinger, 1881)
SUBS Euphydryas aurinia davidi (Oberthür, 1881)

ORIG Melitaea artemis var. davidi Oberthür, 1881
SSSP Melitaea aurinia mandschurica (Staudinger, 1892)

SUBS Euphydryas aurinia debilis Oberthür, 1909

SPEC Euphydryas alexandrina (Staudinger, 1887)

ORIG Melitaea aurinia var. alexandrina Staudinger, 1887
SSYN Eurodryas alexandrina (Staudinger, 1887)

SPEC Euphydryas desfontainii (Godart, 1819)

ORIG Argynnis desfontainii Godart, 1819
SSYN Eurodryas desfontainii (Godart, 1819)

SPEC Euphydryas orientalis (Herrich-Schaeffer, 1845)

ORIG Melitaea orientalis Herrich-Schaeffer, 1845
SSYN Eurodryas orientalis (Herrich-Schaeffer, 1845)

SPEC Euphydryas maturna (Linnaeus, 1758)

ORIG Papilio maturna Linnaeus, 1758
SSYN Papilio agrotera Bergstrasser, 1780
SSYN Melitaea maturna staudingeri Wnukowsky, 1929
SSYN Melitaea maturna uralensis Staudinger, 1871
SSYN Melitaea maturna idunides Fruhstorfer, 1916
SSYN Hypodryas maturna (Linnaeus, 1758)

SPEC Euphydryas intermedia (Ménétriés, 1859)

SSYN Hypodryas intermedia (Ménétriés, 1859)

SUBS Euphydryas intermedia intermedia (Ménétriés, 1859)
SUBS Euphydryas intermedia wolfensbergeri (Frey, 1880)
SUBS Euphydryas intermedia altaica Seitz, 1908
SUBS Euphydryas intermedia mongolica (Staudinger)

SPEC Euphydryas cynthia (Denis & Schiffermüller, 1775)

SSYN Papilio mysia Hübner, 1800
SSYN Melitaea ichnea Boisduval, 1833
SSYN Hypodryas cynthia (Denis & Schiffermüller, 1775)

SPEC Euphydryas iduna (Dalman, 1816)

SSYN Hypodryas iduna (Dalman, 1816)

SUBS Euphydryas iduna iduna (Dalman, 1816)
SUBS Euphydryas iduna alferakyi Korshunov, 1996
SUBS Euphydryas iduna sajana Higgins, 1950

SPEC Euphydryas gillettii (Barnes, 1897)

SSYN Hypodryas gillettii (Barnes, 1897)

SPEC Euphydryas phaeton (Drury, 1773)

ORIG Papilio phaeton Drury, 1773
SSYN Euphydryas phaetaena (Hübner, 1819)
SSYN Euphydryas phaetontea (Godart, 1819)
SSYN Euphydryas superba (Strecker, 1878)
SSYN Euphydryas phaethusa (Hulst, 1881)
SSYN Euphydryas streckeri Ellsworth, 1902
SSYN Euphydryas schausi A. H. Clark, 1927
SSYN Euphydryas magnifica A. H. Clark, 1927
SSYN Euphydryas borealis F. & R. Chermock, 1940

SUBS Euphydryas phaeton phaeton (Drury, 1773)
SUBS Euphydryas phaeton ozarkae Masters, 1968

SPEC Euphydryas editha (Boisduval, 1852)

SSYN Occidryas editha (Boisduval, 1852)

SUBS Euphydryas editha editha (Boisduval, 1852)
SUBS Euphydryas editha alebarki Ferris, 1971
SUBS Euphydryas editha augusta (Edwards, 1890)
SUBS Euphydryas editha aurilacus Gunder, 1928
SUBS Euphydryas editha baroni Edwards, 1879
SUBS Euphydryas editha bayensis Sternitzky, 1937
SUBS Euphydryas editha beani Skinner, 1897
SUBS Euphydryas editha colonia (Wright, 1905)
SUBS Euphydryas editha edithana Strand, 1914
SUBS Euphydryas editha fridayi Gunder, 1931
SUBS Euphydryas editha gunnisonensis Brown, 1971
SUBS Euphydryas editha hutchinsi McDunnough, 1928
SUBS Euphydryas editha insularis Emmel & Emmel, 1974
SUBS Euphydryas editha lawrencei Gunder, 1931
SUBS Euphydryas editha lehmani Gunder, 1929
SUBS Euphydryas editha luestherae Murphy & Ehrlich, 1981
SUBS Euphydryas editha monoensis Gunder, 1928
SUBS Euphydryas editha nubigena Behr, 1863
SUBS Euphydryas editha remingtoni Burdick, 1959
SUBS Euphydryas editha rubicunda (Edwards, 1881)
SUBS Euphydryas editha taylori Edwards, 1888
SUBS Euphydryas editha wrighti Gunder, 1929
SUBS Euphydryas editha quino Behr, 1863

SPEC Euphydryas chalcedona (Doubleday, 1847)

SSYN Occidryas chalcedona (Doubleday, 1847)

SUBS Euphydryas chalcedona chalcedona (Doubleday, 1847)
SUBS Euphydryas chalcedona colon Edwards, 1881
SUBS Euphydryas chalcedona corralensis Emmel & Emmel, 1973
SUBS Euphydryas chalcedona dwinnellei Edwards, 1881
SUBS Euphydryas chalcedona hennei Scott, 1981
SUBS Euphydryas chalcedona huellemanni J.A. Comstock, 1926
SUBS Euphydryas chalcedona kingstonensis Emmel & Emmel, 1973
SUBS Euphydryas chalcedona klotsi dos Passos, 1938
SUBS Euphydryas chalcedona macglashanii Rivers, 1888
SUBS Euphydryas chalcedona olancha Wright, 1905
SUBS Euphydryas chalcedona paradoxa McDunnough, 1927
SUBS Euphydryas chalcedona perdiccas Edwards, 1880
SUBS Euphydryas chalcedona sierra Wright, 1905
SUBS Euphydryas chalcedona sperryi Chermock & Chermock, 1945
SUBS Euphydryas chalcedona wallacensis Gunder, 1928
SUBS Euphydryas chalcedona nevadensis Bauer, 1975

SPEC Euphydryas anicia (Doubleday, 1847)

SSYN Occidryas anicia (Doubleday, 1847)

SUBS Euphydryas anicia anicia (Doubleday, 1847)
SUBS Euphydryas anicia alena Barnes & Benjamin, 1926
SUBS Euphydryas anicia bakeri Stallings & Turner, 1945
SUBS Euphydryas anicia bernadetta Leussler, 1920
SUBS Euphydryas anicia capella Barnes, 1897
SUBS Euphydryas anicia carmentis Barnes & Benjamin, 1926
SUBS Euphydryas anicia chuskae Ferris & R. Holland, 1980
SUBS Euphydryas anicia cloudcrofti Ferris & R. Holland, 1980
SUBS Euphydryas anicia effi Stallings & Turner, 1945
SUBS Euphydryas anicia eurytion Mead, 1875
SUBS Euphydryas anicia helvia Scudder, 1869
SUBS Euphydryas anicia hermosa (Wright, 1905)
SUBS Euphydryas anicia hopfingeri Gunder, 1934
SUBS Euphydryas anicia howlandi Stallings & Turner, 1947
SUBS Euphydryas anicia irelandi Gunder, 1929
SUBS Euphydryas anicia macyi Fender & Jewett, 1953
SUBS Euphydryas anicia magdalena Barnes & McDunnough, 1918
SUBS Euphydryas anicia maria (Skinner, 1899)
SUBS Euphydryas anicia morandi Gunder, 1928
SUBS Euphydryas anicia veazieae Fender & Jewett, 1953
SUBS Euphydryas anicia wheeleri Edwards, 1881
SUBS Euphydryas anicia windi Gunder, 1932

SUBT Melitaeina Newman, 1870
GENU Melitaea Fabricius, 1807

GSYN Schoenis Hübner, 1818
GSYN Cinclidia Hübner, 1818
GSYN Mellicta Billberg, 1820
GSYN Didymaeformia Verity, 1950
GSYN Athaliaeformia Verity, 1950

SPEC Melitaea didyma (Esper, 1779)

ORIG Papilio didyma Esper, 1779
SSYN Papilio cyhteris Mueschen, 1781

SUBS Melitaea didyma didyma (Esper, 1779)
SUBS Melitaea didyma occidentalis Staudinger, 1861
SUBS Melitaea didyma neera Fischer de Waldheim, 1840
SUBS Melitaea didyma turkestanica Sheljuzkho, 1929

SSSP Melitaea pseudoala Sheljuzkho, 1928

SUBS Melitaea didyma mixta Evans, 1932

SPEC Melitaea latonigena Eversmann, 1847
SPEC Melitaea interrupta Kolenati, 1846

SSYN Melitaea transcaucasica Turati, 1919

SPEC Melitaea persea Kollar, 1850

SSYN Melitaea didyma magnacasta Verity, 1929
SSYN Melitaea didyma araratica Verity, 1929
SSYN Melitaea didyma sargon Hemming, 1932
SSYN Melitaea tauricus Belter, 1934

SUBS Melitaea persea persea Kollar, 1850
SUBS Melitaea persea caucasica Staudinger, 1861
SUBS Melitaea persea dodgsoni Grose-Smith, 1887
SUBS Melitaea persea paphlagonia Fruhstorfer, 1917
SUBS Melitaea persea montium Belter, 1934

SPEC Melitaea saxatilis Christoph, 1873
SPEC Melitaea deserticola Oberthür, 1876

SUBS Melitaea deserticola deserticola Oberthür, 1876
SUBS Melitaea deserticola macromaculata Belter, 1934

SPEC Melitaea abyssinica Oberthür, 1909

SUBS Melitaea abyssinica abyssinica Oberthür, 1909
SUBS Melitaea abyssinica scotti Higgins, 1941

SPEC Melitaea casta Kollar, 1849

SUBS Melitaea casta casta Kollar, 1849
SUBS Melitaea casta wiltshirei Higgins, 1941

SPEC Melitaea didymina Staudinger, 1895
SPEC Melitaea didymoides Eversmann, 1847

SUBS Melitaea didymoides didymoides Eversmann, 1847

SSSP Melitaea didyma latonia Grum-Grshmailo, 1891
SSSP Melitaea didyma mandchurica Seitz, 1908
SSSP Melitaea didyma eupatides Fruhstorfer, 1916
SSSP Melitaea didyma seitzi Matsumara, 1929

SUBS Melitaea didymoides pekinensis Seitz, 1909

SPEC Melitaea ala Staudinger, 1881

SUBS Melitaea ala ala Staudinger, 1881

SSSP Melitaea ala allah Bryk, 1940

SUBS Melitaea ala bicolor Seitz, 1907

SPEC Melitaea kotshubeji Sheljuzhko, 1928

SUBS Melitaea kotshubeji kotshubeji Sheljuzhko, 1928
SUBS Melitaea kotshubeji bundeli Kolesnichenko, 1999
SUBS Melitaea kotshubeji kugarti Kolesnichenko, 1999

SPEC Melitaea ninae Sheljuzhko, 1935

SSYN Melitaea rosea Higgins, 1938
SSYN Melitaea ala ella-claudia Bryk, 1940

SPEC Melitaea chitralensis Moore, 1901

SSYN Melitaea didyma chitralipulvia Verity, 1929

SPEC Melitaea enarea Fruhstorfer, 1916

SUBS Melitaea enarea enarea Fruhstorfer, 1916

SSSP Melitaea didyma shugnana Sheljuzhko, 1929

SUBS Melitaea enarea ishkashima Sheljuzhko, 1929
SUBS Melitaea enarea permuta Higgins, 1941
SUBS Melitaea enarea gromenkoi Kolesnichenko, 1999

SPEC Melitaea sutschana Staudinger, 1892
SPEC Melitaea acreina Staudinger, 1886
SPEC Melitaea lunulata Staudinger, 1901
SPEC Melitaea fergana Staudinger, 1882

SSYN Melitaea fergana jacobsoni Higgins, 1941

SPEC Melitaea infernalis Groum-Grshmailo, 1891
SPEC Melitaea ambrisia Higgins, 1935

SUBS Melitaea ambrisia ambrisia Higgins, 1935
SUBS Melitaea ambrisia alraschid Higgins, 1941

SPEC Melitaea shandura Evans, 1924
SPEC Melitaea athene Staudinger, 1881
SPEC Melitaea trivia (Denis & Schiffermüller, 1775)

ORIG Papilio trivia Denis & Schiffermüller, 1775
SSYN Papilio iphigenia Esper, 1782
SSYN Papilio fascelis Esper, 1784
SSYN Papilio antigonus Herbst, 1800
SSYN Melitaea didyma aabaca Fruhstorfer, 1917
SSYN Melitaea catapeloides Stauder, 1918
SSYN Melitaea trivia interrupta Skala, 1907
SSYN Melitaea trivia lathon Fruhstorfer, 1917
SSYN Melitaea robertsi Butler, 1880
SSYN Melitaea trivia tigris Higgins, 1941

SPEC Melitaea yuenty Oberthür, 1886
SPEC Melitaea lutko Evans, 1932

SUBS Melitaea lutko lutko Evans, 1932
SUBS Melitaea lutko mimetica Higgins, 1940

SPEC Melitaea romanovi Groum-Grshmailo, 1891
SPEC Melitaea arduinna (Esper, 1784)

ORIG Papilio arduinna Esper, 1784

SPEC Melitaea avinovi Sheljuzhko, 1914
SPEC Melitaea agar Oberthür, 1886

SSYN Melitaea didyma wardi Watkins, 1927
SSYN Melitaea didyma baileyi Watkins, 1927

SPEC Melitaea cinxia (Linnaeus, 1758)

ORIG Papilio cinxia Linnaeus, 1758
SSYN Papilio pilodellae Rottemburg, 1775
SSYN Papilio delia Schiffermüller, 1775
SSYN Papilio cinxia var. fulla Quesnel, 1791
SSYN Papilio abacus Retzius, 1783

SUBS Melitaea cinxia cinxia (Linnaeus, 1758
SUBS Melitaea cinxia clarissa Staudinger, 1901

SSSP Melitaea cinxia australis Verity, 1916
SSSP Melitaea cinxia narbonensis Fruhstorfer, 1917
SSSP Melitaea cinxia sicula Ragusa, 1920
SSSP Melitaea cinxia deva Hemming, 1934
SSSP Melitaea cinxia castiliana Turati, 1920
SSSP Melitaea cinxia carpetana Verity, 1929

SUBS Melitaea cinxia tschujaca Seitz, 1909

SSSP Melitaea cinxia arelatia Fruhstorfer, 1910

SUBS Melitaea cinxia amardea Groum-Grshmailo, 1895
SUBS Melitaea cinxia atlantis Le Cerf, 1928

SSSP Melitaea cinxia algerica Oberthür, 1914 (nec Melitaea aetherie algerica Heyne, 1893
SSSP Melitaea cinxia eupompe Hemming, 1933

SPEC Melitaea minerva Staudinger, 1881
SPEC Melitaea danieli Achtelik, 1999
SPEC Melitaea sebastiani Achtelik, 1999
SPEC Melitaea gabrielae Achtelik, 1999
SPEC Melitaea fumarata Achtelik, 1999
SPEC Melitaea balba Evans, 1912
SPEC Melitaea balbina Tytler, 1926
SPEC Melitaea elisabethae Avinov, 1910
SPEC Melitaea sultanensis Staudinger, 1886
SPEC Melitaea pallas Staudinger, 1886
SPEC Melitaea turanica Erschoff, 1874

SSYN Melitaea pamira Staudinger, 1887
SSYN Melitaea expressa Groum-Grshmailo, 1887

SPEC Melitaea asteroida Staudinger, 1881

SSYN Melitaea asteroida clara Staudinger, 1887
SSYN Melitaea asteroida uitasica Wagner, 1913

SPEC Melitaea arcesia Bremer, 1864

SSYN Melitaea baikalensis Bremer, 1864
SSYN Melitaea arcesia chuana Groum-Grshmailo, 1893
SSYN Melitaea arcesia carmana Fruhstorfer, 1915
SSYN Melitaea arcesia rucephala Fruhstorfer, 1915

SUBS Melitaea arcesia arcesia Bremer, 1864
SUBS Melitaea arcesia minor Elwes, 1899
SUBS Melitaea arcesia jezabel Oberthür, 1896
SUBS Melitaea arcesia leechi Alpheraki, 1895
SUBS Melitaea arcesia bellona Leech, 1892
SUBS Melitaea arcesia sikkimensis (Moore, 1901)
SUBS Melitaea arcesia sindura Moore, 1865
SUBS Melitaea arcesia amoenula Felder, 1867
SUBS Melitaea arcesia balbita Moore, 1874
SUBS Melitaea arcesia irma Higgins, 1941

SPEC Melitaea diamina (Lang, 1789)

ORIG Papilio diamina Lang, 1789
SSYN Papilio dictynna Esper, 1777
SSYN Papilio hebe Borkhausen, 1793
SSYN Papilio corythalia Hübner, 1800
SSYN Cinclidia orthia Hübner, 1818
SSYN Melitaea dictynna alpestris Fruhstorfer, 1917

SUBS Melitaea diamina diamina (Lang, 1789)
SUBS Melitaea diamina protomedia Ménétriés, 1858

SSSP Melitaea protomedia regama Fruhstorfer, 1915

SPEC Melitaea phoebe (Denis & Schiffermüller, 1775)

ORIG Papilio phoebe Denis & Schiffermüller, 1775
SSYN Papilio corythallia Esper, 1780
SSYN Papilio paedotrophos Bergstrasser, 1780
SSYN Papilio tremulae Piller & Mitterpacher, 1783
SSYN Melitaea phoebe aethereaeformis Verity, 1919
SSYN Melitaea phoebe alatauica Wagner, 1913
SSYN Melitaea phoebe alternans Seitz, 1909
SSYN Melitaea phoebe amanica Rebel, 1917
SSYN Melitaea phoebe canellina Stauder, 1922
SSYN Melitaea phoebe caucasicola Verity, 1919
SSYN Melitaea phoebe changaica Seitz, 1909
SSYN Melitaea phoebe dorae Graves, 1925
SSYN Melitaea phoebe gerinia Fruhstorfer, 1916
SSYN Melitaea phoebe koios Fruhstorfer, 1908
SSYN Melitaea phoebe lokris Fruhstorfer, 1908
SSYN Melitaea phoebe mandarina Seitz, 1908
SSYN Melitaea phoebe minoa Fruhstorfer, 1917
SSYN Melitaea phoebe ogygia Fruhstorfer, 1908
SSYN Melitaea phoebe ottonis Fruhstorfer, 1916
SSYN Melitaea phoebe rovia Fruhstorfer, 1919
SSYN Melitaea phoebe telona Fruhstorfer, 1907
SSYN Melitaea phoebe tungana Seitz, 1909
SSYN Melitaea phoebe tusca Verity, 1919
SSYN Melitaea phoebe wagneri Wnokowsky, 1929

SUBS Melitaea phoebe phoebe (Denis & Schiffermüller, 1775)
SUBS Melitaea phoebe occitanica Staudinger, 1871

SPEC Melitaea scotosia Butler, 1878

SUBS Melitaea scotosia scotosia Butler, 1878
SUBS Melitaea scotosia butleri Higgins, 1940

SPEC Melitaea punica Oberthür, 1876
SPEC Melitaea sarvistana Wiltshire, 1941
SPEC Melitaea aetherie (Hübner, 1826)

ORIG Papilio aetherie Hübner, 1826

SUBS Melitaea aetherie aetherie (Hübner, 1826)
SUBS Melitaea aetherie algerica Ruhl, 1892

SPEC Melitaea sibina Alpheraki, 1881
SPEC Melitaea collina Lederer, 1861
SPEC Melitaea turkmanica Higgins, 1940
SPEC Melitaea consulis Wiltshire, 1941
SPEC Melitaea gina Higgins, 1941
SPEC Melitaea athalia (Rottemburg, 1775)

ORIG Papilio athalia Rottemburg, 1775
SSYN Mellicta athalia (Rottemburg, 1775)

SUBS Melitaea athalia athalia (Rottemburg, 1775)

SSSP Melitaea athalia anatolica Wagner, 1929

SUBS Melitaea athalia celadussa Fruhstorfer, 1910

SSSP Melitaea pseudathalia Reverdin, 1921

SUBS Melitaea athalia nevadensis Oberthür, 1904
SUBS Melitaea athalia norvegica Aurivillius, 1888

SSSP Melitaea athalia scandinavica Staudinger, 1892
SSSP Melitaea athalia lachares Fruhstorfer, 1919
SSSP Melitaea athalia polaris Rygge, 1921
SSSP Melitaea athalia borealis Lindström, 1929

SUBS Melitaea athalia caucasogenita Verity, 1930

SPEC Melitaea ambigua Ménétriés, 1859

ORIG Melitaea athalia var. ambigua Ménétriés, 1859
SSYN Melitaea niphona Butler, 1878
SSYN Melitaea athalia kenteana Seitz, 1909
SSYN Melitaea athalia bathilda Fruhstorfer, 1919
SSYN Melitaea athalia nossis Fruhstorfer, 1919
SSYN Melitaea ambigua sachalinensis Matsumura, 1925
SSYN Melitaea ambigua sayanskalpina Verity, 1940
SSYN Mellicta ambigua (Ménétriés, 1859)

SPEC Melitaea britomartis Assmann, 1847

SSYN Mellicta britomartis (Assmann, 1847)

SUBS Melitaea britomartis britomartis Assmann, 1847

SSSP Melitaea veronicae Dorfmeister, 1853
SSSP Melitaea aurelia marussia Fruhstorfer, 1919

SUBS Melitaea britomartis aureliaeformis (Verity, 1916)

SSSP Melitaea melathalia Rocci, 1930
SSSP Melitaea melathalia taurinorum Rocci, 1932

SUBS Melitaea britomartis amurensis (Staudinger, 1892)

SSSP Melitaea aurelia seminigra Seitz, 1909
SSSP Melitaea britomartis frigidaltaica Verity, 1940

SUBS Melitaea britomartis latefascia (Fixsen, 1883)

SSSP Melitaea coreae Verity, 1930

SPEC Melitaea deione (Geyer, 1832)

ORIG Papilio deione Geyer, 1832
SSYN Mellicta deione (Geyer, 1832)

SUBS Melitaea deione deione (Geyer, 1832)
SUBS Melitaea deione berisalii Ruhl, 1891
SUBS Melitaea deione nitida Oberthür, 1909

SPEC Melitaea centralasiae Wnukowsky, 1929

SSYN Mellicta centralasiae (Wnukowsky, 1929)

SPEC Melitaea aurelia Nickerl, 1850

SSYN Papilio athalia minor Esper, 1784
SSYN Papilio parthenie Borkhausen, 1788 (nec Papilio parthenie Bergstrasser, 1780)
SSYN Mellicta aurelia (Nickerl, 1850)

SUBS Melitaea aurelia aurelia Nickerl, 1850
SUBS Melitaea aurelia albimacula Wojlusiak & Niesiowski, 1946
SUBS Melitaea aurelia distans Higgins, 1955

SPEC Melitaea parthenoides Keferstein, 1851

SSYN Mellicta parthenoides (Keferstein, 1851)

SPEC Melitaea varia Meyer-Dür, 1851

SSYN Mellicta varia (Meyer-Dür, 1851)

SPEC Melitaea asteria (Freyer, 1828)

ORIG Papilio asteria Freyer, 1828
SSYN Mellicta asteria (Freyer, 1828)

SPEC Melitaea menetriesi Caradja, 1895

SSYN Mellicta menetriesi (Caradja, 1895)

SPEC Melitaea rebeli Wnukowsky, 1929

SSYN Melitaea asteria var. altaica Staudinger, 1901 (nec Melitaea didyma altaica Groum-Grshmailo, 1893)
SSYN Mellicta rebeli (Wnukowsky, 1929)

SPEC Melitaea plotina Bremer, 1861

SSYN Melitaea pacifica Verity, 1932
SSYN Mellicta plotina (Bremer, 1861)

SPEC Melitaea alatauica Staudinger, 1881

SSYN Mellicta alatauica (Staudinger, 1881)

SUBT Chlosynina (not available as yet, from Kons 2000)
GENU Chlosyne Butler, 1870

GSYN Limnaecia Scudder, 1872
GSYN Charidryas Scudder, 1872
GSYN Thessalia Scudder, 1875

SPEC Chlosyne palla (Boisduval, 1852)

ORIG Melitaea palla Boisduval, 1852
SSYN Melitaea sterope Edwards, 1870
SSYN Melitaea palla eremiata Wright, 1905
SSYN Melitaea flavula Barnes & McDunnough, 1918
SSYN Melitaea calydon Holland, 1931
SSYN Melitaea hewesi Leussler, 1931
SSYN Chlosyne palla altasierra Emmel, Emmel & Mattoon, 1998
SSYN Chlosyne palla australomontana Emmel, Emmel & Mattoon, 1998

SPEC Chlosyne acastus (W.H. Edwards, 1874)

ORIG Melitaea acastus Edwards, 1874
SSYN Melitaea neumoegeni Skinner, 1895
SSYN Melitaea neumoegeni sabina Wright, 1905
SSYN Melitaea palla vallismortis Johnson, 1938
SSYN Chlosyne acastus waucoba Emmel, Emmel & Mattoon, 1998
SSYN Chlosyne acastus robusta Austin, 1998

SPEC Chlosyne whitneyi (Behr, 1863)

ORIG Melitaea whitneyi Behr, 1863
SSYN Melitaea malcolmi Comstock, 1926

SUBS Chlosyne whitneyi whitneyi (Behr, 1863)
SUBS Chlosyne whitneyi damoetas (Skinner, 1902)

SPEC Chlosyne gabbii (Behr, 1863)

ORIG Melitaea gabbii Behr, 1863
SSYN Melitaea sonorae Boisduval, 1869
SSYN Chlosyne gabbii atrifasciata Hawks & Emmel, 1998

SPEC Chlosyne hoffmanni (Behr, 1863)

ORIG Melitaea hoffmanni Behr, 1863
SSYN Melitaea hellicta Boisduval, 1869
SSYN Melitaea hoffmanni segregata Barnes & MacDunnough, 1918
SSYN Melitaea bridgei Comstock, 1924
SSYN Chlosyne hoffmanni manchada Bauer, 1959

SPEC Chlosyne harrisii (Scudder, 1862)

ORIG Phyciodes harrisii Scudder, 1862
SSYN Phyciodes hanhami Fletcher, 1904
SSYN Melitaea harrisii albimontana Avinoff, 1930
SSYN Melitaea harrisii liggetti Avinoff, 1930

SPEC Chlosyne kendallorum Opler, 1999
SPEC Chlosyne nycteis (Boisduval & LeConte, 1833)

SUBS Chlosyne nycteis nycteis (Doubleday & Hewitson, 1847)

SSSP Melitaea oenone Scudder, 1862

SUBS Chlosyne nycteis drusius (Edwards, 1884)
SUBS Chlosyne nycteis reversa (Chermock & Chermock, 1940)
SUBS Chlosyne nycteis obsoleta Gatrelle, 2003

SPEC Chlosyne gorgone (Hübner, 1810)

ORIG Dryas gorgone Hübner, 1810

SUBS Chlosyne gorgone gorgone (Hübner, 1810)
SUBS Chlosyne gorgone carlota (Reakirt, 1866)

NOMEN DUBIUM Melitaea ismeria Boisduval & LeConte, 1833 See TILS-Leps-talk archives for active discussion of this name in November-December 2003
SPEC Chlosyne lacinia Geyer, 1837

SSYN Synchloe saundersi Doubleday, 1848
SSYN Synchloe tellias Bates, 1864
SSYN Synchloe quehtala Reakirt, 1866
SSYN Synchloe ardema Reakirt, 1866
SSYN Synchloe mediatrix Felder & Felder, 1867
SSYN Synchloe paupera Felder & Felder, 1867
SSYN Synchloe misera Felder & Felder, 1869
SSYN Synchloe pretona Boisduval, 1870
SSYN Synchloe crocale Edwards, 1874
SSYN Chlosyne adjutrix Scudder, 1875
SSYN Synchloe adelina Staudinger, 1875
SSYN Synchloe nigrescens Edwards, 1893
SSYN Synchloe rufescens Edwards, 1893
SSYN Chlosyne lacinia indigens Hall, 1924

SPEC Chlosyne californica (Wright, 1905)

ORIG Synchloe californica Wright, 1905

SPEC Chlosyne definita (Aaron, 1884)

ORIG Melitaea definita Aaron, 1884
SSYN Melitaea albiplaga Aaron, 1884
SSYN Melitaea schausi Godman & Salvin, 1901
SSYN Melitaea beckeri Godman & Salvin, 1901
SSYN Melitaea anastasia Hemming, 1934

SPEC Chlosyne ezra (Hewitson, 1864)

ORIG Eresia ezra Hewitson, 1864
SSYN Eresia yorita Reakirt, 1865

SPEC Chlosyne chinatiensis (Tinkham, 1944)

ORIG Melitaea chinatiensis Tinkham, 1944

SPEC Chlosyne perlula (Felder & Felder, 1861)

ORIG Eresia perlula Felder & Felder, 1861
SSYN Phyciodes hondana Weymer, 1890

SPEC Chlosyne theona (Ménétriés, 1855)

ORIG Melitaea theona Ménétriés, 1855
SSYN Melitaea thekla Edwards, 1870
SSYN Melitaea bolli Edwards, 1877
SSYN Thessalia theona costaricensis Austin & Smith, 1998
SSYN Thessalia theona brocki Austin & Smith, 1998
SSYN Thessalia theona mullinsi Austin & Smith, 1998
SSYN Thessalia theona minimus Austin & Smith, 1998

SPEC Chlosyne leanira (Felder & Felder, 1860)

ORIG Melitaea leanira Felder & Felder, 1860

SUBS Chlosyne leanira leanira (Felder & Felder, 1860)

SSSP Melitaea leanira obsoleta Edwards, 1877
SSSP Melitaea leanira obliterata Strecker, 1878
SSSP Melitaea alma Strecker, 1878
SSSP Melitaea wrightii Edwards, 1886
SSSP Melitaea cerrita Wright, 1905
SSSP Melitaea leona Wright, 1905
SSSP Lemonias alma koebeli Gunder, 1927
SSSP Melitaea leanira daviesi Wind, 1947
SSSP Thessalia leanira oregonensis Bauer, 1975
SSSP Thessalia leanira elegans Priestaf & Emmel, 1998
SSSP Thessalia leanira nebularum Austin & Smith, 1998
SSSP Thessalia leanira austrima Austin & Smith, 1998
SSSP Thessalia leanira basinensis Austin & Smith, 1998
SSSP Thessalia leanira flavodorsalis Austin & Smith, 1998

SUBS Chlosyne leanira fulvia Edwards, 1879

SSSP Thessalia fulvia paraiensis Smith & Brock, 1988
SSSP Thessalia fulvia coronado Smith & Brock, 1988

SUBS Chlosyne leanira cyneas (Godman & Salvin, 1878)
SUBS Chlosyne leanira cynisca (Godman & Salvin, 1882)

SPEC Chlosyne endeis (Godman & Salvin, 1894)

ORIG Synchloe endeis Godman & Salvin, 1894

SUBS Chlosyne endeis endeis (Godman & Salvin, 1894)
SUBS Chlosyne endeis pardelina Higgins, 1960

SPEC Chlosyne marina (Geyer, 1837)

ORIG Araschnia marina Geyer, 1837

SPEC Chlosyne melitaeoides (Felder & Felder, 1867

ORIG Synchloe melitaeoides Felder & Felder, 1867

SPEC Chlosyne erodyle (Bates, 1864)

ORIG Synchloe erodyle Bates, 1864

SUBS Chlosyne erodyle erodyle (Bates, 1864)
SUBS Chlosyne erodyle poecile (Felder & Felder, 1867)

SPEC Chlosyne melanarge (Bates, 1864)

ORIG Synchloe melanarge Bates, 1864

SPEC Chlosyne eumeda (Godman & Salvin, 1894)

ORIG Synchloe eumeda Godman & Salvin, 1894
SSYN Synchloe dryope Godman & Salvin, 1894
SSYN Chlosyne fasciata Röber, 1914

SPEC Chlosyne hylaeus (Godman & Salvin, 1894

ORIG Synchloe hylaeus Godman & Salvin, 1894

SPEC Chlosyne ehrenbergi (Geyer, 1833)

ORIG Morpheis ehrenbergi Geyer, 1833

SPEC Chlosyne hippodrome (Geyer, 1837)

ORIG Araschnia hippodrome Geyer, 1837

SPEC Chlosyne narva (Fabricius, 1793)

ORIG Papilio narva Fabricius, 1793
SSYN Cethosia bonpland Latreille, 1809

SPEC Chlosyne gaudealis (Bates, 1864)

ORIG Synchloe gaudealis Bates, 1864
SSYN Chlosyne gaudealis laeta Röber, 1914

SUBS Chlosyne gaudealis gaudealis (Bates, 1864)
SUBS Chlosyne gaudealis wellingi Miller & Roger, 1979

SPEC Chlosyne janais (Drury, 1782)

ORIG Papilio janais Drury, 1782
SSYN Chlosyne hyperia marianna Röber, 1914
SSYN Chlosyne hyperia irrubescens Hall, 1924
SSYN Chlosyne gloriosa Bauer, 1959

SPEC Chlosyne rosita Hall, 1924

SUBS Chlosyne rosita rosita Hall, 1924
SUBS Chlosyne rosita browni Bauer, 1961
SUBS Chlosyne rosita montana Hall, 1924
SUBS Chlosyne rosita riobalensis Bauer, 1961
SUBS Chlosyne rosita mazarum Miller & Roger, 1979

GENU Microtia Bates, 1864

GSYN Texola Higgins, 1959
GSYN Dymasia Higgins, 1960

SPEC Microtia elada (Hewitson, 1868)

SUBS Microtia elada elada (Hewitson, 1868)

ORIG Eresia elada Hewitson, 1868
SSYN Melitaea callina Boisduval, 1869
SSYN Eresia socia Felder & Felder, 1869
SSYN Phyciodes hepburni Godman & Salvin, 1901

SUBS Microtia elada perse (Edwards, 1882)

ORIG Melitaea perse Edwards, 1882

SUBS Microtia elada ulrica (Edwards, 1877)

ORIG Melitaea ulrica Edwards, 1877
SSYN Melitaea immitata Strecker, [1878]

SPEC Microtia elva Bates, 1864

SUBS Microtia elva elva Bates, 1864
SUBS Microtia elva horni Rebel, 1906

SSYN Microtia elva f. draudti Röber, [1914]

SPEC Microtia dymas (W.H. Edwards, 1877)

SUBS Microtia dymas dymas (W.H. Edwards, 1877)

ORIG Melitaea dymas Edwards, 1877
SSYN Melitaea larunda Strecker, [1878]
SSYN Melitaea senrabii Barnes, 1900

SUBS Microtia dymas chara (W.H. Edwards, [1884])

ORIG Melitaea chara Edwards, [1884]

SUBS Microtia dymas imperialis Bauer, 1959

SSYN Melitaea chara ab. jacintoi Gunder, 1924
SSYN Melitaea chara ab. nitela J.A. Comstock, 1926

SPEC Microtia anomalus (Godman & Salvin, 1897)

ORIG Phyciodes anomalus Godman & Salvin, 1897

SPEC Microtia coracara (Dyar, 1912)

ORIG Phyciodes coracara Dyar, 1912
SSYN Phyciodes albipunctata Röber, 1913

GENU Poladryas Bauer, 1961

SPEC Poladryas minuta (W.H. Edwards, 1861)

ORIG Melitaea minuta Edwards, 1861
SSYN Melitaea approximata Strecker, 1900

SPEC Poladryas arachne (Edwards, 1869)

SUBS Poladryas arachne arachne (Edwards, 1869)

ORIG Melitaea arachne Edwards, 1869
SSYN Melitaea arachne ab. gunderiae Holland, 1930

SUBS Poladryas arachne gilensis (Holland, 1930)

ORIG Melitaea gilensis Holland, 1930

SUBS Poladryas arachne nympha (Edwards, 1884)

ORIG Melitaea nympha Edwards, 1884
SSYN Melitaea arachne ab. skinneri Oberthür, 1923
SSYN Melitaea pola var. arachne ab. polingi Gunder, 1926

SUBS Poladryas arachne expedita Austin, 1998
SUBS Poladryas arachne monache (Comstock, 1918)

ORIG Melitaea monache Comstock, 1918

GENU Atlantea Higgins, 1959

SPEC Atlantea cryptadia Sommer & Schwartz, 1980
SPEC Atlantea perezi (Herrich-Schäffer, 1862)

ORIG Synchloe perezi Herrich-Schäffer, 1862

SPEC Atlantea tulita (Dewitz, 1877)

ORIG Synchloe tulita Dewitz, 1877

SPEC Atlantea pantoni (Kaye, 1906)

ORIG Chlosyne pantoni Kaye, 1906
SSYN Chlosyne seitzi Röber, 1914

SUBT Gnathotrichina (not available as yet, from Kons 2000)
GENU Gnathotriche Felder & Felder, 1862

GSYN Gnathotrusia Higgins, 1981
SPEC Gnathotriche exclamationis (Kollar, 1850)

SUBS Gnathotriche exclamationis exclamationis (Kollar, 1850)

ORIG Euterpe exclamationis Kollar, 1850
SSYN Eresia eleates Weymer, 1907
SSYN Phyciodes vanessoides Röber, [1914]

SUBS Gnathotriche exclamationis elaea (Hewitson, 1869)

ORIG Eresia elaea Hewitson, 1869

SUBS Gnathotriche exclamationis sodalis Staudinger, 1885

SPEC Gnathotriche mundina (Druce, 1876)

SUBS Gnathotriche mundina mundina (Druce, 1876)

ORIG Eresia mundina Druce, 1876

SUBS Gnathotriche mundina erebia (Röber, [1914])

ORIG Phyciodes erebia Röber, [1914]
SSYN Phyciodes erebia mima Bryk, 1953

SUBS Gnathotriche mundina fallax (Staudinger, 1885)

ORIG Phyciodes fallax Staudinger, 1885
SSYN Phyciodes mundina f. callianira A. Hall, 1930

SUBS Gnathotriche mundina hopfferi (Röber, [1914])

ORIG Phyciodes hopfferi Röber, [1914]
SSYN Phyciodes mundina testacea A. Hall, 1935
SSYN Actinote erinome f. postaperta Bryk, 1950

SUBS Gnathotriche mundina rosina (Dognin, 1888)

ORIG Phyciodes rosina Dognin, 1888
SSYN Phyciodes mundina peraea Hall, 1928

SUBS Gnathotriche mundina steinii (Dewitz, 1877)

ORIG Acraea steinii Dewitz, 1877
SSYN Acraea atrata Dewitz, 1877
SSYN Eresia epione Godman & Salvin, 1878
SSYN Phyciodes styx Staudinger, 1885
SSYN Phyciodes callianthina Hall, 1921
SSYN Phyciodes epione elenae Hall, 1928

GENU Higginsius Hemming, 1964

GSYN Fulvia Higgins, 1959

SPEC Higginsius fasciata (Hoppfer, 1874)

ORIG Melitaea fasciata Hoppfer, 1874
SSYN Eresia pearcei Druce, 1876
SSYN Phyciodes fasciatella Röber, 1913

SPEC Higginsius miriam (Dognin, 1888)

ORIG Phyciodes miriam Dognin, 1888

SUBT Phyciodina Higgins, 1981
GENU Antillea Higgins, 1959

SPEC Antillea pelops (Drury, 1773)

SUBS Antillea pelops pelops (Drury, 1773)

ORIG Papilio pelops Drury, 1773

SUBS Antillea pelops anocaona (Herrich-Schäffer, 1865)

ORIG Melitaea anocaona Herrich-Schäffer, 1865

SUBS Antillea pelops pygmaea (Godart, 1819)

ORIG Argynnis pygmea Godart, 1819
SSYN Papilio aegon Fabricius, 1781
SSYN Erycina aedon Godart, [1824]

SPEC Antillea proclea (Doubleday, [1847])

ORIG Melitaea proclea Doubleday, [1847]

GENU Phystis Higgins, 1981

SPEC Phystis simois (Hewitson, 1864)

SUBS Phystis simois simois (Hewitson, 1864)

ORIG Eresia simois Hewitson, 1864
SSYN Phyciodes pedrona Moulton, 1909

SUBS Phystis simois variegata (Röber, 1913)

ORIG Phyciodes variegata Röber, 1913
NUDU Phyciodes simois var. nigrina Breyer, 1930
SSYN Phyciodes simois ab. nigrina Hayward, 1931

SUBS Phystis simois pratti (A. Hall, 1935)

ORIG Phyciodes simois pratti Hall, 1935

SUBS Phystis simois chinchipensis (Hayward, 1964)

ORIG Phyciodes chinchipensis Hayward, 1964

GENU Mazia Higgins, 1981

SPEC Mazia amazonica (Bates, 1864)

SUBS Mazia amazonica amazonica (H.W. Bates, 1864)

ORIG Melitaea amazonica Bates, 1864

SUBS Mazia amazonica cocha Lamas, 1995
SUBS Mazia amazonica tambopata Lamas, 1995

GENU Phyciodes Hübner, 1819

SPEC Phyciodes graphica (Felder, 1869)

SUBS Phyciodes graphica graphica (Felder, 1869)

ORIG Eresia graphica Felder, 1869
SSYN Phyciodes vesta vestalis A. Hall, 1929

SUBS Phyciodes graphica vesta (W.H. Edwards, 1869)

ORIG Melitaea vesta Edwards, 1869
SSYN Phyciodes boucardi Godman & Salvin, 1878
SSYN Phyciodes vesta f. hiemalis W.H. Edwards, 1878
SSYN Phyciodes vesta f. aestiva W.H. Edwards, 1878

SPEC Phyciodes picta (Edwards, 1865)

SUBS Phyciodes picta picta (Edwards, 1865)

ORIG Melitaea picta Edwards, 1865

SUBS Phyciodes picta canace Edwards, 1871

SSYN Phyciodes picta ab. jemezensis Brehme, 1913

SPEC Phyciodes pallescens (Felder, 1869)

ORIG Eresia pallescens Felder, 1869

SPEC Phyciodes phaon (Edwards, 1864)

ORIG Melitaea phaon Edwards, 1864

SUBS Phyciodes phaon phaon (Edwards, 1864)
SUBS Phyciodes phaon maya
SUBS Phyciodes phaon jalapeno Scott, 1998

SPEC Phyciodes orseis Edwards, 1871

SUBS Phyciodes orseis orseis Edwards, 1871
SUBS Phyciodes orseis herlani Bauer, 1975

SPEC Phyciodes mylitta (Edwards, 1861)

ORIG Melitaea mylitta Edwards, 1861
SSYN Melitaea collina Behr, 1863
SSYN Melitaea epula Boisduval, 1869

SUBS Phyciodes mylitta mylitta (Edwards, 1864)
SUBS Phyciodes mylitta arida (Skinner, 1917)
SUBS Phyciodes mylitta arizonensis Bauer, 1975
SUBS Phyciodes mylitta mexicana Hall, 1928
SUBS Phyciodes mylitta thebais Godman & Salvin, 1878

SPEC Phyciodes pallida (Edwards, 1864)

ORIG Melitaea pallida Edwards, 1864
SSYN Eresia mata Reakirt, 1866

SUBS Phyciodes pallida pallida Edwards, 1864
SUBS Phyciodes pallida barnesi Skinner, 1897

SPEC Phyciodes tharos (Drury, 1773)

ORIG Papilio tharos Drury, 1773
SSYN Papilio euclea Bergstraesser, 1780
SSYN Melitaea marcia Edwards, 1868
SSYN Melitaea packardii Saunders, 1869

SUBS Phyciodes tharos tharos (Drury, 1773)

SSSP Phyciodes tharos distincta Bauer, 1975

SUBS Phyciodes tharos riocolorado Scott, 1992
SUBS Phyciodes tharos orantain Scott, 1998

SPEC Phyciodes cocyta (Cramer, 1777)

ORIG Papilio cocyta Cramer, 1777
SSYN Papilio morpheus Fabricius, 1775
SSYN Phyciodes pascoensis Wright, 1906
SSYN Phyciodes arctica dos Passos, 1935

SUBS Phyciodes cocyta cocyta (Cramer, 1777)
SUBS Phyciodes cocyta selenis (Kirby, 1837)
SUBS Phyciodes cocyta diminutor Scott, 1998
SUBS Phyciodes cocyta arenacolor Austin, 1998

SPEC Phyciodes batesii (Reakirt, 1865)

ORIG Eresia batesii Reakirt, 1865

SUBS Phyciodes batesii batesii (Reakirt, 1865)
SUBS Phyciodes batesii lakota Scott, 1994
SUBS Phyciodes batesii anasazi Scott, 1994
SUBS Phyciodes batesii apsaalooke Scott, 1994
SUBS Phyciodes batesii maconensis Gatrelle, 1998

SPEC Phyciodes pulchella (Boisduval, 1852)

ORIG Melitaea pulchella Boisduval, 1852
SSYN Melitaea campestris Behr, 1863
SSYN Melitaea pratensis Behr, 1863

SUBS Phyciodes pulchella pulchella (Boisduval, 1852)
SUBS Phyciodes pulchella montana (Behr, 1863)

SSSP Melitaea orsa Boisduval, 1869

SUBS Phyciodes pulchella camillus Edwards, 1871

SSSP Phyciodes emissa Edwards, 1871

SUBS Phyciodes pulchella tutchone Scott, 1994
SUBS Phyciodes pulchella shoshoni Scott, 1994
SUBS Phyciodes pulchella owimba Scott, 1998
SUBS Phyciodes pulchella deltarufa Scott, 1998
SUBS Phyciodes pulchella vallis Austin, 1998
SUBS Phyciodes pulchella inornatus Austin, 1998

GENU Tegosa Higgins, 1981

SPEC Tegosa claudina (Eschscholtz, 1821)

ORIG Acraea claudina Eschscholtz, 1821
SSYN Papilio thymetus Fabricius, 1787
SSYN Argynnis flavia Godart, [1824]
SSYN Phyciodes liriope f. claudina ab. immaculata Hayward, 1935
SSYN Tegosa similis Higgins, 1981

SPEC Tegosa orobia (Hewitson, 1864)

SUBS Tegosa orobia orobia (Hewitson, 1864)

ORIG Eresia orobia Hewitson, 1864

SUBS Tegosa orobia ursula (Staudinger, 1894)

ORIG Phyciodes ursula Staudinger, 1894

SPEC Tegosa fragilis (Bates, 1864)

ORIG Melitaea fragilis Bates, 1864

SPEC Tegosa infrequens Higgins, 1981
SPEC Tegosa flavida (Hewitson, 1868)

ORIG Eresia flavida Hewitson, 1868
SSYN Phyciodes albescens Röber, 1913

SPEC Tegosa tissoides (Hall, 1928)

ORIG Phyciodes tissoides Hall, 1928

SPEC Tegosa pastazena (Bates, 1864)

ORIG Melitaea liriope var. pastazena Bates, 1864
SSYN Phyciodes liriope flavina Röber, 1913

SPEC Tegosa guatemalena (Bates, 1864)

ORIG Melitaea fragilis var. guatemalana Bates, 1864

SPEC Tegosa anieta (Hewitson, 1864)

SUBS Tegosa anieta anieta (Hewitson, 1864)

ORIG Eresia anieta Hewitson, 1864

SUBS Tegosa anieta cluvia (Godman & Salvin, 1882)

ORIG Phyciodes cluvia Godman & Salvin, 1882

SUBS Tegosa anieta lirina (Röber, 1913)

ORIG Phyciodes liriope lirina Röber, 1913

SUBS Tegosa anieta luka Higgins, 1981

SPEC Tegosa serpia Higgins, 1981
SPEC Tegosa nazaria (Felder & Felder, 1867)

ORIG Eresia nazaria Felder & Felder, 1867
SSYN Phyciodes aquila Hall, 1917

SPEC Tegosa etia (Hewitson, 1868)

ORIG Eresia etia Hewitson, 1868
SSYN Eresia tissa Hewitson, 1869
SSYN Phyciodes etia selenoides Hall, 1928
SSYN Phyciodes melini Bryk, 1953

SPEC Tegosa nigrella (Bates, 1866)

SUBS Tegosa nigrella nigrella Bates, 1866

ORIG Melitaea nigrella Bates, 1866
SSYN Phyciodes lutescens Godman & Salvin, 1882

SUBS Tegosa nigrella niveonotis (Butler & Druce, 1872)

ORIG Phyciodes niveonotis Butler & Druce, 1872

SPEC Tegosa selene (Röber, 1913)

ORIG Phyciodes selene Röber, 1913

GENU Tisona Higgins, 1981

SPEC Tisona saladillensis (Giacomelli, 1911)

SUBS Tisona saladillensis saladillensis (Giacomelli, 1911)

ORIG Phyciodes saladillensis Giacomelli, 1911

SUBS Tisona saladillensis clarior Higgins, 1981

GENU Anthanassa Scudder, 1875

GSYN Tritanassa Forbes, 1945

SPEC Anthanassa drusilla (Felder & Felder, 1861)

ORIG Eresia drusilla Felder & Felder, 1861
SSYN Phyciodes flavimacula Röber, 1913

SUBS Anthanassa drusilla drusilla (Felder & Felder, 1861)
SUBS Anthanassa drusilla lelex (Bates, 1864)

SSSP Melitaea alethes Bates, 1864
SSSP Melitaea stesilea Bates, 1864

SUBS Anthanassa drusilla alceta (Hewitson, 1869)

SSSP Phyciodes flavimacula conflua Röber, 1913

SUBS Anthanassa drusilla verena (Hewitson, 1864)

SPEC Anthanassa ptolyca (Bates, 1864)

ORIG Melitaea ptolyca Bates, 1864
SSYN Phyciodes carrera Hall, 1917

SUBS Anthanassa ptolyca ptolyca (Bates, 1964)
SUBS Anthanassa ptolyca amator (Hall, 1929)

SPEC Anthanassa ardys (Hewitson, 1864)

ORIG Eresia ardys Hewitson, 1864
SSYN ?Eresia genigueh Reakirt, 1865

SUBS Anthanassa ardys ardys (Hewitson, 1864)
SUBS Anthanassa ardys subota (Godman & Salvin, 1878)

SPEC Anthanassa dracaena (Felder & Felder, 1867)

ORIG Eresia dracaena Felder & Felder, 1867

SPEC Anthanassa phlegias (Godman & Salvin, 1901)

ORIG Phyciodes phlegias Godman & Salvin, 1901
SSYN Phyciodes carigia Schaus, 1902
SSYN Phyciodes platytaenia Röber, 1913

SPEC Anthanassa texana (Edwards, 1863)

ORIG Melitaea texana Edwards, 1863
SSYN Eresia smerdis Hewitson, 1864

SUBS Anthanassa texana texana (Edwards, 1863)
SUBS Anthanassa texana seminole (Skinner, 1911)

SPEC Anthanassa alexon (Godman & Salvin, 1889)

ORIG Phyciodes alexon Godman & Salvin, 1889
SSYN ?Phyciodes natacles Dyar, 1913

SUBS Anthanassa alexon alexon (Godman & Salvin, 1889)
SUBS Anthanassa alexon subconcolor (Röber, 1913)

SPEC Anthanassa acesas (Hewitson, 1864)

ORIG Eresia acesas Hewitson, 1864
SSYN ?Phyciodes annita Staudinger, 1885

SPEC Anthanassa nebulosa (Godman & Salvin, 1878)

ORIG Phyciodes nebulosa Godman & Salvin, 1878
SSYN Phyciodes castianira Godman & Salvin, 1880

SPEC Anthanassa argentea (Godman & Salvin, 1882)

ORIG Phyciodes argentea Godman & Salvin, 1882

SPEC Anthanassa atronia (Bates, 1866)

ORIG Melitaea atronia Bates, 1866
SSYN ?Eresia sydra Reakirt, 1866
SSYN Eresia obscurata Felder, 1869
SSYN Phyciodes diallus Godman & Salvin, 1878
SSYN ?Phyciodes chromis Godman & Salvin, 1878
SSYN Phyciodes cassiopea Godman & Salvin, 1878
SSYN ?Phyciodes albofascia Röber, 1913

SPEC Anthanassa otanes (Hewitson, 1864)

ORIG Eresia otanes Hewitson, 1864

SUBS Anthanassa otanes otanes (Hewitson, 1864)
SUBS Anthanassa otanes sopolis (Godman & Salvin, 1878)

SSSP Phyciodes cyno Godman & Salvin, 1889
SSSP Phyciodes dora Schaus, 1913

SPEC Anthanassa annulata Higgins, 1981
SPEC Anthanassa crithona (Salvin, 1871)

ORIG Melitaea crithona Salvin, 1871

SPEC Anthanassa fulviplaga (Butler, 1872)

ORIG Phyciodes fulviplaga Butler, 1872
SSYN Phyciodes crithona stenotaenia Röber, 1913

SPEC Anthanassa hermas (Hewitson, 1864)

ORIG Eresia hermas Hewitson, 1864
SSYN Eresia conferta Felder & Felder, 1867
SSYN ?Phyciodes brancodia Schaus, 1902
SSYN Phyciodes aequatorialis Röber, 1913

SPEC Anthanassa frisia (Poey, 1832)

ORIG Melitaea frisia Poey, 1832
SSYN Eresia gyges Hewitson, 1864

SPEC Anthanassa tulcis (Bates, 1864)

ORIG Melitaea tulcis Bates, 1864
SSYN ?Eresia archesilea Felder, 1869
SSYN Eresia punctata Edwards, 1871

SPEC Anthanassa dubia (Hall, 1929)

ORIG Phyciodes frisia dubia Hall, 1929

SPEC Anthanassa taeniata (Röber, 1913)

ORIG Phyciodes taeniata Röber, 1913

SPEC Anthanassa sosis (Godman & Salvin, 1878)

ORIG Phyciodes sosis Godman & Salvin, 1878

SPEC Anthanassa drymaea (Godman & Salvin, 1878)

ORIG Phyciodes drymea Godman & Salvin, 1878

SPEC Anthanassa sitalces (Godman & Salvin, 1882)

ORIG Phyciodes sitalces Godman & Salvin, 1882
SSYN Phyciodes sitalces saltator Hall, 1929

SPEC Anthanassa cortes (Hall, 1917)

ORIG Phyciodes cortes Hall, 1917

GENU Dagon Higgins, 1981

SPEC Dagon catula (Hopffer, 1874)

ORIG Eresia catula Hopffer, 1874
SSYN Phyciodes birivula Dyar, 1913
SSYN Phyciodes minima Röber, 1913
SSYN Phyciodes catula extincta Röber, 1913
NUDU Phyciodes abasina Röber, 1913
SSYN Phyciodes catula ab. fulvocincta (A. Hall, 1929)
SSYN Phyciodes catagraphus Hayward, 1933

SPEC Dagon pusilla (Salvin, 1869)

ORIG Eresia pusilla Salvin, 1869
SSYN Eresia calena Hopffer, 1874
SSYN Phyciodes adoxa Erschoff, 1874

SPEC Dagon morena (Röber, 1913)

ORIG Phyciodes morena Röber, 1913

GENU Telenassa Higgins, 1981

SPEC Telenassa abas (Hewitson, 1864)

ORIG Eresia abas Hewitson, 1864

SPEC Telenassa berenice (C. Felder & R. Felder, 1862)

SUBS Telenassa berenice berenice (C. Felder & R. Felder, 1862)

ORIG Eresia berenice C. Felder & R. Felder, 1862
NUDU Phyciodes telemachus Röber, 1913
SSYN Phyciodes teletusa douglasi (Bryk, 1953)

SUBS Telenassa berenice drusinilla (Röber, 1913)

ORIG Phyciodes drusinilla Röber, 1913
SSYN Phyciodes teletusa signata A. Hall, 1928

SPEC Telenassa delphia (C. Felder & R. Felder, 1861)

SUBS Telenassa delphia delphia (C. Felder & R. Felder, 1861)

ORIG Eresia delphia C. Felder & R. Felder, 1861
SSYN Eresia aceta Hewitson, 1864
SSYN Eresia ezba Hewitson, 1868

SUBS Telenassa delphia catenaria (Godman & Salvin, 1880)

ORIG Phyciodes catenaria Godman & Salvin, 1880

SUBS Telenassa delphia elaphina (Röber, 1913)

ORIG Phyciodes elaphina Röber, 1913

SUBS Telenassa delphia flavocincta (Dognin, 1887)

ORIG Phyciodes flavocincta Dognin, 1887

SUBS Telenassa delphia gaujoni (Dognin, 1887)

ORIG Phyciodes gaujoni Dognin, 1887

SUBS Telenassa delphia nana (H. Druce, 1874)

ORIG Phyciodes nana H. Druce, 1874
SSYN Eresia geminia Hopffer, 1874
SSYN Phycoides [sic] omosis (Dyar, 1913)

SUBS Telenassa delphia nussia (H. Druce, 1876)

ORIG Eresia nussia H. Druce, 1876

SUBS Telenassa delphia trimaculata (Hewitson, 1869)

ORIG Eresia trimaculata Hewitson, 1869
SSYN Phyciodes taphius Godman & Salvin, 1878
SSYN Phyciodes elaphiaea abrupta Röber, 1913

SPEC Telenassa fontus (A. Hall, 1928)

SUBS Telenassa fontus fontus (A. Hall, 1928)

ORIG Phyciodes fontus A. Hall, 1928
SSYN Phyciodes rima A. Hall, 1929

SUBS Telenassa fontus schmitzorum (Austin & Mielke, 1993)

ORIG Castilia schmitzorum Austin & Mielke, 1993

SPEC Telenassa jana (C. Felder & R. Felder, 1867)

ORIG Eresia jana C. Felder & R. Felder, 1867
SSYN Eresia elaphiaea Hewitson, 1868

SPEC Telenassa notus (A. Hall, 1917)

ORIG Phyciodes notus A. Hall, 1917
SSYN Phyciodes notus f. pullopicta A. Hall, 1929

SPEC Telenassa sepultus (A. Hall, 1928)

ORIG Phyciodes sepultus A. Hall, 1928

SPEC Telenassa teletusa (Godart, [1824])

SUBS Telenassa teletusa teletusa (Godart, [1824])

ORIG Argynnis teletusa Godart, [1824]
SSYN Eresia comaela Reakirt, 1866
SSYN Phyciodes teletusa boliviana Röber, 1913

SUBS Telenassa teletusa burchelli (Moulton, 1909)

ORIG Phyciodes burchelli Moulton, 1909
SSYN Phyciodes teletusa peruana Röber, 1913

GENU Ortilia Higgins, 1981

SPEC Ortilia liriope (Cramer, 1775)

ORIG Papilio liriope Cramer, 1775

SPEC Ortilia gentina Higgins, 1981
SPEC Ortilia orthia (Hewitson, 1864)

ORIG Eresia orthia Hewitson, 1864
SSYN Phyciodes poltis Godman & Salvin, 1878
SSYN Phyciodes orthia ab. evanescens Röber, 1913
SSYN Phyciodes orthia röberi [sic] A. Hall, 1935

SPEC Ortilia orticas (Schaus, 1902)

SUBS Ortilia orticas orticas (Schaus, 1902)

ORIG Phyciodes orticas Schaus, 1902

SUBS Ortilia orticas zamora (Hall, 1917)

ORIG Phyciodes zamora Hall, 1917

SPEC Ortilia sejona (Schaus, 1902)

ORIG Phyciodes sejona Schaus, 1902

SPEC Ortilia velica (Hewitson, 1864)

SUBS Eresia velica velica (Hewitson, 1864)

ORIG Eresia velica Hewitson, 1864

SUBS Eresia velica durnfordi (Godman & Salvin, 1878)

ORIG Phyciodes durnfordi Godman & Salvin, 1878
SSYN Phyciodes velica dictynna Röber, 1913

SPEC Ortilia dicoma (Hewitson, 1864)

ORIG Eresia dicoma Hewitson, 1864

SPEC Ortilia polinella (Hall, 1928)

ORIG Phyciodes polinella Hall, 1928

SPEC Ortilia ithra (Kirby, 1900)

ORIG Phyciodes ithra Kirby, 1900
NUDU Neptis ithra Kirby, 1871
SSYN Phyciodes ianthe atra Röber, 1913
SSYN Phyciodes janthe ab. nigra Giacomelli, 1928
SSYN Phyciodes ithra rufocincta Hall, 1928
SSYN Phyciodes janthe ab. schaeferi Köhler, 1945
SSYN Ortilia ithra luiseana Anken & Silva, 1998

GENU Eresia Boisduval, 1836

GYSN Neptis Illiger, 1807 suppressed

SPEC Eresia clio (Linnaeus, 1758)

ORIG Papilio clio Linnaeus, 1758
SSYN Eresia clara Bates, 1864
SSYN Phyciodes clio estebana Hall, 1929
SSYN Phyciodes clio reducta Hall, 1929
SPEC Eresia nauplius (Linnaeus, 1758)

ORIG Papilio nauplius Linnaeus, 1758

SUBS Eresia nauplius nauplius (Linnaeus, 1758)
SUBS Eresia nauplius extensa (Hall, 1929)

SPEC Eresia plagiata (Röber, 1913)

ORIG Phyciodes nauplia plagiata Röber, 1913

SPEC Eresia letitia Hewitson, 1869

SSYN ?Phyciodes leucophaea Weymer, 1890

SUBS Eresia letitia letitia Hewitson, 1869
SUBS Eresia letitia ocellata (Röber, 1913)

SSSP Eresia neptoides Rosenberg & Talbot, 1914
SSSP Eresia letitia nigra Rosenberg & Talbot, 1914

SPEC Eresia lansdorfi (Godart, 1819)

ORIG Heliconia lansdorfi Godart, 1819
SSYN Eresia lansdorfi f. jacinthica (Röber, 1913)
SSYN Eresia lansdorfi f. veternosa (Ferreira d'Almeida, 1922)
SSYN Eresia lansdorfi f. sulphurata (Zikán, 1937)

SPEC Eresia sestia Hewitson, 1869

SSYN Phyciodes saturata Röber, 1913

SPEC Eresia coela Druce, 1874
SPEC Eresia oblita (Staudinger, 1885)

ORIG Phyciodes oblita Staudinger, 1885

SPEC Eresia carme Doubleday, 1847

SUBS Eresia carme carme Doubleday, 1847
SUBS Eresia carme laias Godman & Salvin, 1879

SSSP Phyciodes laias lycus Hall, 1928

SPEC Eresia polina Hewitson, 1852

SSYN Eresia encina Felder & Felder, 1852
SSYN Phyciodes polina f. intermedia Röber, 1913

SPEC Eresia alsina Hewitson, 1869
SPEC Eresia cissia (Hall, 1928)

ORIG Phyciodes ithomioides cissia Hall, 1928

SPEC Eresia eutropia Hewitson, 1874
SPEC Eresia mimas (Staudinger, 1885)

ORIG Phyciodes mimas Staudinger, 1885

SPEC Eresia quintilla Hewitson, 1872
SPEC Eresia poecilina Bates, 1866

SSYN Phyciodes eutropia confirmans Hall, 1930

SPEC Eresia melaina Higgins, 1981
SPEC Eresia sticta Schaus, 1913
SPEC Eresia ithomioides Hewitson, 1864

SUBS Eresia ithomioides ithomioides Hewitson, 1864
SUBS Eresia ithomioides pseudocelemina (Strand,1916)

SPEC Eresia anomala Higgins, 1981
SPEC Eresia nigripennis Salvin, 1869

SSYN Eresia dismorphina Butler, 1872

SPEC Eresia emerantia Hewitson, 1857

SSYN Phyciodes celemina Röber, 1913

SPEC Eresia moesta Salvin & Godman, 1868

SSYN Eresia ildica Hewitson, 1869
SSYN Eresia cerquita Dognin, 1894
SSYN Phyciodes ildica fassli Röber, 1913

SPEC Eresia phaedima Salvin & Godman, 1868

SSYN Eresia prisca Hopffer, 1874
SSYN Melitaea magniplaga Röber, 1913

SPEC Eresia datis Hewitson, 1864

SUBS Eresia datis datis Hewitson, 1864

SSSP ?Phyciodes fenestrata Röber, 1913

SUBS Eresia datis corybassa Hewitson, 1874

SSSP Phyciodes cornelia Staudinger, in litt.; Röber, 1914
SSSP ?Phyciodes corybassa mimicry Bryk, 1953

SPEC Eresia margaretha Hewitson, 1872

SSYN Phyciodes margaretha manto Hall, 1930

SPEC Eresia eunice (Hübner, 1807)

ORIG Nereis fulva eunice Hübner, 1807
SSYN Eresia pella Hewitson, 1852

SUBS Eresia eunice eunice (Hübner, 1807)
SUBS Eresia eunice olivancia Bates, 1864

SSSP Eresia drypetis Godman & Salvin, 1878
SSSP Eresia klagesii Weeks, 1906
SSSP Eresia olivencia f. gudruna Röber, 1913
SSSP Eresia olivencia f. brunhilda Röber, 1913
SSSP Eresia olivencia polymnia Röber, 1913
SSSP ?Phyciodes eunice homogena Bryk, 1953

SUBS Eresia eunice esora Hewitson, 1857

SPEC Eresia etesiae (Hall, 1929)

ORIG Phyciodes erysice etesiae Hall, 1929

SPEC Eresia erysice (Geyer, 1832)

ORIG Melinaea erysice Geyer, 1832

SPEC Eresia casiphia Hewitson, 1869
SPEC Eresia mechanitis Godman & Salvin, 1878
SPEC Eresia pelonia Hewitson, 1852

SSYN Eresia ithomiola Salvin, 1869
SSYN Phyciodes callonia Staudinger, 1885
SSYN Phyciodes callonia var. murena Staudinger, 1885
SSYN Eresia callonioides Strand, 1912
SSYN Phyciodes pardalina Röber, 1913
SSYN Phyciodes pardalina apicalis Röber, 1913
SSYN Phyciodes murena heliconina Röber, 1913

SPEC Eresia phillyra Hewitson, 1852

SSYN Eresia ezorias Hewitson, 1852

SPEC Eresia aveyrona Bates, 1864

SUBS Eresia aveyrona aveyrona Bates, 1864
SUBS Eresia aveyrona mylitta Hewitson, 1869

SSSP Eresia bella Hewitson, 1869
SSSP Phyciodes bella Kirby, 1871

SPEC Eresia perna Hewitson, 1852

SSYN Phyciodes alma Staudinger, 1885

SPEC Eresia levina Hewitson, 1872

SSYN Morpheis felderi Hopffer, 1874
SSYN ?Phyciodes levina werneri Herin, 1925

SPEC Eresia actinote Salvin, 1869

SSYN Eresia acraea Hopffer, 1874
SSYN Phyciodes actinote limbata Röber, 1913

GENU Castilia Higgins, 1981

SPEC Castilia castilla (Felder & Felder, 1862)

SUBS Castilia castilla castilla (Felder & Felder, 1862)

ORIG Eresia castilla Felder & Felder, 1862
SSYN Phyciodes castilla f. virilis Röber, [1914]

SUBS Castilia castilla occidentalis (Fassl, 1912)

ORIG Eresia castilla f. occidentalis Fassl, 1912

SPEC Castilia perilla (Hewitson, 1852)

ORIG Eresia perilla Hewitson, 1852
SSYN Eresia acraeina Hewitson, [1864]
SSYN Eresia aricilla Hopffer, 1874
SSYN Eresia heliconoides Butler, 1877
SSYN Phyciodes acraeina f. hilarina Röber, 1913
SSYN Eresia neria f. microdryope Röber, 1913
SSYN Eresia amoenides Röber, 1913
SSYN Phyciodes acraeina f. aberrans Röber, 1913
SSYN Phyciodes perilla lugubris Röber, 1914
SSYN Phyciodes pellenea Röber, 1914
SSYN Phyciodes pellenea crucifera Bryk, 1953

SPEC Castilia chiapaensis (Beutelspacher, 1990)

ORIG Anthanassa chiapaensis Beutelspacher, 1990

SPEC Castilia neria (Hewitson, 1869)

ORIG Eresia neria Hewitson, 1869
SSYN Phyciodes aurora Röber, 1914

SPEC Castilia nortbrundii (Weeks, 1901)

ORIG Phyciodes nortbrundii Weeks, 1901
SSYN Eresia nebrites Weymer, 1907
SSYN Phyciodes actinotina Röber, 1914

SPEC Castilia eranites (Hewitson, 1857)

ORIG Eresia eranites Hewitson, 1857
SSYN Phyciodes eranites mejicana Röber, 1913

SPEC Castilia fulgora (Godman & Salvin, 1878)

ORIG Phyciodes fulgora Godman & Salvin, 1878
SSYN Phyciodes levana Röber, 1913
SSYN Phyciodes faustus Godman & Salvin, 1897

SPEC Castilia ofella (Hewitson, 1864)

ORIG Eresia ofella Hewitson, 1864

SPEC Castilia myia (Hewitson, 1874)

ORIG Eresia myia Hewitson, 1874

SPEC Castilia griseobasalis (Röber, 1913)

ORIG Phyciodes myia f. griseobasalis Röber, 1913

SPEC Castilia angusta (Hewitson, 1868)

ORIG Eresia angusta Hewitson, 1868
SSYN Castilia longala Austin & T.C. Emmel, 1996

SPEC Castilia chinantlensis (de la Maza, 1978)

ORIG Phyciodes chinantlensis de la Maza, 1978

SPEC Castilia guaya (A. Hall, 1929)

ORIG Phyciodes guaya A. Hall, 1929

GENU Janatella Higgins, 1981

SPEC Janatella leucodesma (Felder & Felder, 1861)

ORIG Eresia leucodesma Felder & Felder, 1861
SSYN Eresia cincta Edwards, 1864

SPEC Janatella hera (Cramer, 1779)

ORIG Papilio hera Cramer, 1779
SSYN Papilio ianthe Fabricius, 1781

SPEC Janatella fellula (Schaus, 1902)

ORIG Phyciodes fellula Schaus, 1902

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