Apaturinae

Niklas Wahlberg

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Updated 2023-04-23. Based on

Markku Savela's pages

Friedlander, T. P. 1988. Taxonomy, phylogeny, and biogeography of Asterocampa Röber 1916 (Lepidoptera, Nymphalidae, Apaturinae). Journal for Research on the Lepidoptera 25: 215-338.

The Butterflies of America

SUBF Apaturinae

GENU Apatura Fabricius, 1807

GSYN Potamis Hübner, 1806
GSYN Apaturia Sodoffsky, 1837
GSYN Athymodes Moore, 1896

SPEC Apatura iris (Linnaeus, 1758)

ORIG Papilio iris Linnaeus, 1758

SUBS Apatura iris iris (Linnaeus, 1758)
SUBS Apatura iris bieti Oberthür, 1885
SUBS Apatura iris xanthina Oberthür, 1909
SUBS Apatura iris amurensis Stichel, 1908
SUBS Apatura iris kansuensis O. Bang-Haas, 1933
SUBS Apatura iris peninsularis Lee & Takamura, 1981

SPEC Apatura ilia (Schiffermüller, 1775)

ORIG Papilio ilia Schiffermüller, 1775

SUBS Apatura ilia ilia (Schiffermüller, 1775)
SUBS Apatura ilia praeclara Bollow, 1930
SUBS Apatura ilia here Felder & Felder, 1862
SUBS Apatura ilia sobrina Stichel
SUBS Apatura ilia serarum Oberthür
SUBS Apatura ilia yunnana Mell, 1952
SUBS Apatura ilia phaedra Leech, 1892
SUBS Apatura ilia hereoides O. Bang-Haas, 1933
SUBS Apatura ilia herastituta O. Bang-Haas, 1936
SUBS Apatura ilia pusilla O. Bang-Haas, 1936
SUBS Apatura ilia szechwanensis Le Moult, 1947
SUBS Apatura ilia extensa Le Moult, 1947
SUBS Apatura ilia subsobrina Mell, 1952
SUBS Apatura ilia huapingensis Yoshino, 1998

SPEC Apatura metis Freyer, 1829

SSYN Apatura ilia krylovi Kurentzov, 1937
SSYN Apatura iris abramovi Kurentzov, 1970

SUBS Apatura metis metis Freyer, 1829
SUBS Apatura metis bunea Herrich-Schäffer, 1845
SUBS Apatura metis substituta Butler, 1873
SUBS Apatura metis irtyshika Korshunov, 1982
SUBS Apatura metis heijona Matsumura, 1928
SUBS Apatura metis doii Matsumura, 1928
SUBS Apatura metis separata Tuzov, 2000

SPEC Apatura laverna Leech, 1893

SUBS Apatura laverna laverna Leech, 1893

SSYN Apatura laverna leii Chou, 1994

SUBS Apatura laverna yunlingensis Yoshino, 1999

GENU Mimathyma Moore, 1896

GSYN Bremeria Moore, 1896
GSYN Amuriana Korshunov & Dubatolov, 1984

SPEC Mimathyma schrenckii (Ménétriés, 1859)

SUBS Apatura schrenckii schrenckii (Ménétriés, 1859)
SUBS Apatura schrenckii laeta Oberthür, 1906

SPEC Mimathyma nycteis (Ménétriés, 1859)

ORIG Atyma [sic] nycteis Ménétriés, 1859
SSYN Atyma [sic] cassiope Ménétriés, 1859

SPEC Mimathyma chevana (Moore, 1866)

ORIG Athyma chevana Moore, 1866

SPEC Mimathyma ambica (Kollar, 1844)

SUBS Apatura ambica ambica (Kollar, 1844)
SUBS Apatura ambica namouna (Doubleday, 1845)
SUBS Apatura ambica garlanda Fruhstorfer, 1913
SUBS Apatura ambica miranda (Fruhstorfer, 1902)
SUBS Apatura ambica claribella (Fruhstorfer, 1902)
SUBS Apatura ambica martini (Fruhstorfer, 1906)

GENU Chitoria Moore, 1896

SPEC Chitoria sordida (Moore, 1866)

ORIG Apatura sordida Moore, 1866

SUBS Chitoria sordida sordida (Moore, 1866)
SUBS Chitoria sordida vietnamica Nguyen, 1979

SPEC Chitoria modesta (Oberthür, 1906)
SPEC Chitoria naga (Tytler, 1915)
SPEC Chitoria ulupi (Doherty, 1889)

SUBS Chitoria ulupi ulupi (Doherty, 1889)
SUBS Chitoria ulupi dubernardi (Oberthür)
SUBS Chitoria ulupi arakii Naritomi, 1959
SUBS Chitoria ulupi kalaurica (Tytler, 1926)
SUBS Chitoria ulupi fulva (Leech, 1891)
SUBS Chitoria ulupi morii (Seok, 1937)
SUBS Chitoria ulupi esakii Shirôzu, 1959
SUBS Chitoria ulupi setia (Fruhstorfer, 1909)
SUBS Chitoria ulupi tong Yoshino, 1997

SPEC Chitoria chrysolora (Fruhstorfer, 1908)
SPEC Chitoria subcaerulea (Leech, 1891)

ORIG Apatura subcaerulea Leech, 1891

SPEC Chitoria cooperi (Tytler, 1926)
SPEC Chitoria fasciola (Leech, 1890)

ORIG Apatura fasciola Leech, 1890

GENU Lelecella Hemming, 1939

GSYN Lelex de Nicéville, 1900 (preocc. Lelex Rafinesque, 1815)

SPEC Lelecella limenitoides (Oberthür, 1890)

ORIG Vanessa limenitoides Oberthür, 1890
SSYN Lelex limenitoides (Oberthür, 1890)

SUBS Lelex limenitoides limenitoides (Oberthür, 1890)
SUBS Lelex limenitoides wangi Chou, 1994

GENU Dilipa Moore, 1857

SPEC Dilipa fenestra (Leech, 1891)
SPEC Dilipa morgiana (Westwood, 1850)

ORIG Apatura morgiana Westwood, 1850

GENU Euapatura Ebert, 1971

GSYN Protapatura Shirôzu & Saigusa, 1971

SPEC Euapatura mirza Ebert, 1971

SSYN Protapatura iwasei Igarashi, 1971

GENU Eulaceura Butler, 1872

SPEC Eulaceura manipurensis Tytler, 1915
SPEC Eulaceura osteria (Westwood, 1850)

ORIG Apatura osteria Westwood, 1850

SUBS Eulaceura osteria osteria (Westwood, 1850)
SUBS Eulaceura osteria kumana Fruhstorfer, 1913
SUBS Eulaceura osteria nicomedeia Fruhstorfer
SUBS Eulaceura osteria jembala Fruhstorfer
SUBS Eulaceura osteria bipupillata Lathy
SUBS Eulaceura osteria sitarama Fruhstorfer

GENU Rohana Moore, 1880

SPEC Rohana parisatis (Westwood, 1850)

ORIG Apatura parisatis Westwood, 1850

SUBS Rohana parisatis parisatis (Westwood, 1850)
SUBS Rohana parisatis camiba Moore, 1880
SUBS Rohana parisatis atacinius Fruhstorfer
SUBS Rohana parisatis pseudosiamensis Nguyen-Phung, 1985
SUBS Rohana parisatis staurakius (Fruhstorfer, 1913)
SUBS Rohana parisatis hainana (Fruhstorfer)
SUBS Rohana parisatis sumatrensis Staudinger
SUBS Rohana parisatis javana Fruhstorver, 1906
SUBS Rohana parisatis borneana Fruhstorfer, 1905
SUBS Rohana parisatis nana Staudinger

SPEC Rohana tonkiniana Fruhstorfer, 1906

SUBS Rohana tonkiniana tonkiniana Fruhstorfer, 1906
SUBS Rohana tonkiniana siamensis (Fruhstorfer, 1913)

SPEC Rohana nakula (Moore, 1858)

SUBS Rohana nakula nakula (Moore, 1858)
SUBS Rohana nakula artaxes (de Nicéville, 1895)
SUBS Rohana nakula bernardii Nguyen-Phung, 1985
SUBS Rohana nakula thantoana Kimura, 1994
SUBS Rohana nakula panna Yoshino, 1995

SPEC Rohana ruficincta Lathy, 1913
SPEC Rohana rhea (Felder & Felder, 1863)

SUBS Rohana rhea rhea (Felder & Felder, 1863)
SUBS Rohana rhea danae Fruhstorfer, 1906
SUBS Rohana rhea mindora Fruhstorfer, 1906
SUBS Rohana rhea rana Staudinger

SPEC Rohana macar Wallace, 1869

SUBS Rohana macar macar Wallace, 1869
SUBS Rohana macar pagenstecheri de Nicéville
SUBS Rohana macar butongensis Tsukada, 1991

SPEC Rohana parvata (Moore, 1857)

SUBS Rohana parvata parvata (Moore, 1857)
SUBS Rohana parvata burmana (Tytler, 1940)

GENU Euripus Doubleday, 1848

GSYN Idrusia Corbet, 1943

SPEC Euripus consimilis (Westwood, 1850)

SUBS Euripus consimilis consimilis (Westwood, 1850)
SUBS Euripus consimilis meridionalis Wood-Mason, 1881
SUBS Euripus consimilis eurinus Fruhstorfer, 1903

SPEC Euripus nyctelius (Doubleday, 1845)

ORIG Diadema nyctelius Doubleday, 1845
SSYN Euripus halitherses Doubleday, 1848

SUBS Euripus nyctelius nyctelius (Doubleday, 1845)
SUBS Euripus nyctelius mastor Fruhstorfer, 1903
SUBS Euripus nyctelius euploeoides Felder & Felder, [1867]
SUBS Euripus nyctelius niasicus Fruhstorfer, 1899
SUBS Euripus nyctelius javanus Fruhstorfer, 1899
SUBS Euripus nyctelius borneensis Distant & Pryer, 1887
SUBS Euripus nyctelius palawanicus Fruhstorfer, 1899
SUBS Euripus nyctelius clytia Felder & Felder, [1867]
SUBS Euripus nyctelius orestheion Fruhstorfer, 1914
SUBS Euripus nyctelius nysia Semper, 1887
SUBS Euripus nyctelius sumatrensis Fruhstorfer, 1914
SUBS Euripus nyctelius ophelion Fruhstorfer, 1914

SPEC Euripus robustus Wallace, 1869

GENU Helcyra Felder, 1860

SPEC Helcyra chionippe Felder, 1860

SUBS Helcyra chionippe chionippe Felder, 1860
SUBS Helcyra chionippe thyiada Fruhstorfer, 1913
SUBS Helcyra chionippe kibleri Weymer, 1913
SUBS Helcyra chionippe marginata Rothschild & Jordan, 1899

SPEC Helcyra superba Leech, 1890

SUBS Helcyra superba superba Leech, 1890
SUBS Helcyra superba takamukui Matsumura
SUBS Helcyra superba wiensis Yoshino, 1997

SPEC Helcyra hemina Hewitson, 1864

SUBS Helcyra hemina hemina Hewitson, 1864
SUBS Helcyra hemina masinia Fruhstorfer, 1903

SPEC Helcyra plesseni (Fruhstorfer, 1913)
SPEC Helcyra miyamotoi Koiwaya, 2003
SPEC Helcyra subalba (Poujade, 1885)

SUBS Helcyra subalba subalba (Poujade, 1885)
SUBS Helcyra subalba subsplendens (Mell)

SPEC Helcyra celebensis Martin, 1913

SUBS Helcyra celebensis celebensis Martin, 1913
SUBS Helcyra celebensis fabulose Tsukada, 1991
SUBS Helcyra celebensis semifusca Masui, 1999

GENU Herona Doubleday, 1848

SPEC Herona marathus Doubleday, 1848

SUBS Herona marathus marathus Doubleday, 1848
SUBS Herona marathus angustata Moore, 1878
SUBS Herona marathus andamana Moore, 1877
SUBS Herona marathus marathon Fruhstorfer, 1906
SUBS Herona marathus stellaris Tsukada, 1991

SPEC Herona sumatrana Moore, 1881

SUBS Herona sumatrana sumatrana Moore, 1881
SUBS Herona sumatrana dusuntua Corbet, 1937
SUBS Herona sumatrana schoenbergi Staudinger, 1890
SUBS Herona sumatrana djarang Fruhstorfer, 1893
SUBS Herona sumatrana pringondani Fruhstorfer, 1893

GENU Hestina Westwood, 1850

GSYN Diagora Snellen, 1894
GSYN Hestinalis Bryk, 1938

SPEC Hestina nama (Doubleday, 1844)

ORIG Diadema nama Doubleday, 1844

SUBS Hestina nama nama (Doubleday, 1844)
SUBS Hestina nama namida Fruhstorfer, 1906
SUBS Hestina nama melanoides Joicey & Talbot, 1921

SPEC Hestina mimetica Butler, 1874

SUBS Hestina mimetica mimetica Butler, 1874
SUBS Hestina mimetica ruvanella Fruhstorfer, 1913
SUBS Hestina mimetica carolinae Snellen

SPEC Hestina divona (Hewitson, 1861)
SPEC Hestina dissimilis Hall, 1935
SPEC Hestina waterstradti Watkins, 1928
SPEC Hestina japonica (Felder & Felder, 1862)

ORIG Apatura japonica Felder & Felder, 1862
SSYN Diadema diagoras Hewitson, 1863

SUBS Hestina japonica japonica
SUBS Hestina japonica seoki Shirôzu, 1955

SPEC Hestina assimilis (Linnaeus, 1758)

ORIG Papilio assimilis Linnaeus, 1758
SSYN Hestina nigrivena Leech, 1890

SUBS Hestina assimilis assimilis (Linnaeus, 1758)
SUBS Hestina assimilis formosana Moore, 1896
SUBS Hestina assimilis shirakii Shirôzu, 1955

SPEC Hestina mena Moore, 1858

SUBS Hestina mena mena Moore, 1858
SUBS Hestina mena subdecora (Fruhstorfer, 1903)

SPEC Hestina persimilis (Westwood, 1850)

SUBS Hestina persimilis persimilis (Westwood, 1850)
SUBS Hestina persimilis zella Butler, 1869
SUBS Hestina persimilis chinensis (Leech, 1890)

SPEC Hestina nicevillei Moore, 1895

SUBS Hestina nicevillei nicevillei Moore, 1895
SUBS Hestina nicevillei jermyni (Druce, 1911)
SUBS Hestina nicevillei ouvradi Riley, 1939
SUBS Hestina nicevillei nigra Morishita, 1997
SUBS Hestina nicevillei magna Omoto & Funahashi, 2004

SPEC Hestina namoides de Nicéville, 1900

GENU Sasakia Moore, 1896

SPEC Sasakia charonda (Hewitson, 1863)

ORIG Diadema charonda Hewitson, 1863

SUBS Sasakia charonda charonda (Hewitson, 1863)
SUBS Sasakia charonda coreana (Leech, 1887)
SUBS Sasakia charonda yunnanensis Fruhstorfer
SUBS Sasakia charonda formosana Shirôzu, 1963

SPEC Sasakia funebris (Leech, 1891)

SUBS Sasakia funebris funebris (Leech, 1891)
SUBS Sasakia funebris genestieri Oberthür
SUBS Sasakia funebris fulguralis Matsumura

SPEC Sasakia pulcherrima Chou & Li

GENU Sephisa Moore, 1882

GSYN Castalia Westwood, 1850 (preocc. Castalia Lamarck, 1810, Castalia Savigny, 1822, Castalia Laporte & Gory, 1837)

SPEC Sephisa dichroa (Kollar, 1844)

ORIG Limenitis dichroa Kollar, 1844

SPEC Sephisa princeps (Fixsen, 1887)

SUBS Sephisa princeps princeps (Fixsen, 1887)
SUBS Sephisa princeps tamla Sugiyama, 1999

SPEC Sephisa chandra (Moore, 1858)

ORIG Castalia chandra Moore, 1858

SUBS Sephisa chandra chandra (Moore, 1858)
SUBS Sephisa chandra stubbsi Corbet, 1941
SUBS Sephisa chandra androdamas Fruhstorfer, 1908
SUBS Sephisa chandra zhejiangana Tong, 1994

SPEC Sephisa daimio Matsumura, 1910

GENU Apaturina Herrich-Schaeffer, 1864

SPEC Apaturina erminea (Cramer, 1779)

ORIG Papilio erminea Cramer, 1779
SSYN Apaturina aruensis Joicey & Talbot, 1924
SSYN Apaturina splendidissima Bryk, 1939
SSYN Apaturina principalis Bryk, 1939
SSYN Apaturina androtropa Bryk, 1939

SUBS Apaturina erminea erminea (Cramer, 1779)
SUBS Apaturina erminea mirona Fruhstorfer, 1904
SUBS Apaturina erminea erinna Fruhstorfer, 1904
SUBS Apaturina erminea ribbei Röber, 1886
SUBS Apaturina erminea octavia Fruhstorfer, 1904
SUBS Apaturina erminea papuana Ribbe, 1886

SSYN Apaturina microps Röber, 1894

SUBS Apaturina erminea antonia Fruhstorfer, 1904
SUBS Apaturina erminea xanthocera Rothschild, 1904

SSYN Apaturina kietana Niepelt, 1916

SUBS Apaturina erminea aluna Fruhstorfer, 1904
SUBS Apaturina erminea neopommerania Hagen, 1897

GENU Apaturopsis Aurivillius, 1894

SPEC Apaturopsis cleochares (Hewitson, 1873)

ORIG Apatura cleochares Hewitson, 1873

SUBS Apaturopsis cleochares cleochares (Hewitson, 1873)
SUBS Apaturopsis cleochares schultzei Schmidt, 1921

SPEC Apaturopsis kilusa Grose-Smith, 1891
SPEC Apaturopsis paulianii Viette, 1962

GENU Timelaea Lucas, 1883

SPEC Timelaea maculata (Bremer & Grey, 1852)

ORIG Melitaea? maculata Bremer & Grey, 1852

SPEC Timelaea albescens (Oberthür, 1886)

SUBS Timelaea albescens albescens (Oberthür, 1886)
SUBS Timelaea albescens formosana Fruhstorfer, 1908
SUBS Timelaea albescens longshenensis Yoshino, 1998

SPEC Timelaea aformis Chou, 1994
SPEC Timelaea radiata Chou & Wang, 1994
SPEC Timelaea nana Leech, 1893

GENU Thaleropis Staudinger, 1871

SPEC Thaleropis ionia (Fischer de Waldheim & Eversmann, 1851)

ORIG Vanessa ionia Fischer de Waldheim & Eversmann, 1851

GENU Asterocampa Röber, 1916

GSYN Celtiphaga Barnes & Lindsey, 1922

SPEC Asterocampa celtis (Boisduval & Leconte, 1834)

ORIG Apatura celtis Boisduval & Leconte, 1834

SUBS Asterocampa celtis celtis (Boisduval & Leconte, 1834)

SSYN Apatura alicia Edwards, 1868
SSYN Apatura alba Wright, 1905

SUBS Asterocampa celtis antonia (Edwards, 1878)

SSYN Apatura montis Edwards, 1883

SPEC Asterocampa reinthali Friedlander, 1987
SPEC Asterocampa leilia (Edwards, 1874)

ORIG Apatura leilia Edwards, 1874
SSYN Apatura cocles Edwards, 1884

SPEC Asterocampa clyton (Boisduval & Le Conte, 1833)

ORIG Apatura clyton (Boisduval & Le Conte, 1833)

SUBS Asterocampa clyton clyton (Boisduval & LeConte, 1833)

SSYN Apatura ocellata Edwards, 1876
SSYN Apatura proserpina Scudder, 1868
SSYN Apatura nig Smith, 1903

SUBS Asterocampa clyton texana (Skinner, 1911)

SSYN Apatura subpallida (Barnes & McDunnough, 1913)

SUBS Asterocampa clyton flora (Edwards, 1876)
SUBS Asterocampa clyton louisa Stallings & Turner, 1947

SPEC Asterocampa idyja (Geyer, 1828)

ORIG Doxocopa idyja Geyer, 1828

SUBS Asterocampa idyja idyja Geyer, 1828

SSYN Apatura padola Fruhstorfer, 1912

SUBS Asterocampa idyja argus Bates, 1864

GENU Doxocopa Hübner, 1819

GSYN Catargyria Hübner, 1823
GSYN Chlorippe Doubleday, 1844

SPEC Doxocopa agathina (Cramer, 1777)

SUBS Doxocopa agathina agathina (Cramer, 1777)
SUBS Doxocopa agathina vacuna (Godart, [1824])

SPEC Doxocopa callianira (Ménétriés, 1855)
SPEC Doxocopa clothilda (C. Felder & R. Felder, 1867)
SPEC Doxocopa cyane (Latreille, [1813])

SUBS Doxocopa cyane cyane (Latreille, [1813])
SUBS Doxocopa cyane burmeisteri (Godman & Salvin, 1884)
SUBS Doxocopa cyane mexicana Bryk, 1953
SUBS Doxocopa cyane vespertina Lamas, 1999

SPEC Doxocopa elis (C. Felder & R. Felder, 1861)
SPEC Doxocopa excelsa (Gillott, 1927)
SPEC Doxocopa kallina (Staudinger, 1886)
SPEC Doxocopa laure (Drury, 1773)

SUBS Doxocopa laure laure (Drury, 1773)
SUBS Doxocopa laure druryi (Hübner, [1825])
SUBS Doxocopa laure griseldis (C. Felder & R. Felder, 1862)
SUBS Doxocopa laure laura (Hübner, [1823])
SUBS Doxocopa laure laurona (Schaus, 1902)
SUBS Doxocopa laure mima (Fruhstorfer, 1907)

SPEC Doxocopa laurentia (Godart, [1824])

SUBS Doxocopa laurentia laurentia (Godart, [1824])
SUBS Doxocopa laurentia cherubina (C. Felder & R. Felder, 1867)
SUBS Doxocopa laurentia thalysia (Fruhstorfer, 1907)

SPEC Doxocopa lavinia (A. Butler, 1866)
SPEC Doxocopa linda (C. Felder & R. Felder, 1862)

SUBS Doxocopa linda linda (C. Felder & R. Felder, 1862)
SUBS Doxocopa linda carwa Lamas, 1999
SUBS Doxocopa linda godmani (Dannatt, 1904)
SUBS Doxocopa linda mileta (Boisduval, 1870)
SUBS Doxocopa linda plesaurina (A. Butler & H. Druce, 1872)

SPEC Doxocopa pavon (Latreille, [1809])

SUBS Doxocopa pavon pavon (Latreille, [1809])
SUBS Doxocopa pavon theodora (Lucas, 1857)

SPEC Doxocopa thoe (Godart, [1824])
SPEC Doxocopa zalmunna (A. Butler, 1869)
SPEC Doxocopa zunilda (Godart, [1824])

SUBS Doxocopa zunilda zunilda (Godart, [1824])
SUBS Doxocopa zunilda felderi (Godman & Salvin, 1884)
SUBS Doxocopa zunilda floris (Fruhstorfer, 1907)

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