Anaeini

Niklas Wahlberg

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Updated 2023-07-07. Based on

• The Butterflies of America
• Dias, F. M. S., Janzen, D., Hallwachs, W., Chacón, I., Willmott, K., Ortiz-Acevedo, E., Mielke, O. H. H. & Casagrande, M. M. 2018: DNA barcodes uncover hidden taxonomic diversity behind the variable wing patterns in the Neotropical butterfly genus Zaretis (Lepidoptera: Nymphalidae: Charaxinae). Zoological Journal of the Linnean Society doi:10.1093/zoolinnean/zly036
• Dias, F. M. S., Riley, T. J., Casagrande, M. M. & Mielke, O. H. H. 2021: An overview of “arginussa” species group of Memphis Hübner, [1819], with the description of a new species from Panama (Lepidoptera: Nymphalidae: Charaxinae). Zootaxa 5061: 249–270 doi:10.11646/zootaxa.5061.2.2
• Núñez, R., Genaro, J. A., Miller, J. Y., Barro-Cañamero, A., Álvarez, Y., Fernández, D. M., Minno, M. C., Turner, T., Turlan, V., Wahlberg, N., Pérez-Asso, A. R. & Hausmann, A. 2023: DNA barcodes of the Greater Antillean butterflies (Lepidoptera: Papilionoidea) suggest a richer, more isolated fauna and higher endemism. Biological Journal of the Linnean Society, in press. doi:10.1093/biolinnean/blad015

TRIB Anaeini

GENU Coenophlebia C. Felder & R. Felder, 1862

SPEC Coenophlebia archidona (Hewitson, 1860)

GENU Consul Hübner, [1807]

GSYN Protogonius Hübner, 1819

SPEC Consul electra (Westwood, 1850)

SUBS Consul electra electra (Westwood, 1850)
SUBS Consul electra adustus Lamas, 2003

SPEC Consul excellens (H. Bates, 1864)

SUBS Consul excellens excellens (H. Bates, 1864)
SUBS Consul excellens genini (Le Cerf, 1922)

SPEC Consul fabius (Cramer, 1776)

SUBS Consul fabius fabius (Cramer, 1776)
SUBS Consul fabius albinotatus (A. Butler, 1874)
SUBS Consul fabius bogotanus (A. Butler, 1874)
SUBS Consul fabius castaneus (A. Butler, 1874)
SUBS Consul fabius cecrops (E. Doubleday, [1849])
SUBS Consul fabius diffusus (A. Butler, 1875)
SUBS Consul fabius divisus (A. Butler, 1874)
SUBS Consul fabius drurii (A. Butler, 1874)
SUBS Consul fabius fassli (Röber, 1916)
SUBS Consul fabius fulvus (A. Butler, 1875)
SUBS Consul fabius ochraceus (A. Butler, 1874)
SUBS Consul fabius quadridentatus (A. Butler, 1874)
SUBS Consul fabius semifulvus (A. Butler, 1875)
SUBS Consul fabius superba (Niepelt, 1923)

SPEC Consul panariste (Hewitson, 1856)

SUBS Consul panariste panariste (Hewitson, 1856)
SUBS Consul panariste jansoni (Salvin, 1871)
SUBS Consul panariste ludmilla (Fassl, 1912)
SUBS Consul panariste pandrosa (Niepelt, 1927)

GENU Hypna Hübner, [1819]

SPEC Hypna clytemnestra (Cramer, 1777)

SUBS Hypna clytemnestra clytemnestra (Cramer, 1777)
SUBS Hypna clytemnestra corumbaensis Talbot, 1928
SUBS Hypna clytemnestra forbesi Godman & Salvin, 1884
SUBS Hypna clytemnestra huebneri A. Butler, 1866
SUBS Hypna clytemnestra mexicana A. Hall, 1917
SUBS Hypna clytemnestra negra C. Felder & R. Felder, 1862
SUBS Hypna clytemnestra rufescens A. Butler, 1866

SPEC Hypna iphigenia Herrich-Schäffer, 1862

GENU Polygrapha Staudinger, 1887

SPEC Polygrapha cyanea (Salvin & Godman, 1868)

SUBS Polygrapha cyanea cyanea (Salvin & Godman, 1868)
SUBS Polygrapha cyanea silvaorum Constantino & Salazar, 1998

SPEC Polygrapha suprema (Schaus, 1920)
SPEC Polygrapha tyrianthina (Salvin & Godman, 1868)
SPEC Polygrapha xenocrates (Westwood, 1850)

SUBS Polygrapha xenocrates xenocrates (Westwood, 1850)
SUBS Polygrapha xenocrates ernestoi (Salazar & Constantino, 2001)
SUBS Polygrapha xenocrates punctimarginale (Kaye, 1918)

GENU Siderone Hübner, [1823]

SPEC Siderone galanthis (Cramer, 1775)

SUBS Siderone galanthis nemesis (Illiger, 1801)
SUBS Siderone galanthis galanthis (Cramer, 1775)
SUBS Siderone galanthis thebais C. Felder & R. Felder, 1862

SPEC Siderone syntyche Hewitson, [1854]

SUBS Siderone syntyche syntyche Hewitson, [1854]
SUBS Siderone syntyche angustifascia A. Hall, 1917
SUBS Siderone syntyche mars H. Bates, 1860

GENU Zaretis Hübner, [1819]

SPEC Zaretis itys (Cramer, 1777)
SPEC Zaretis isidora (Cramer, 1779)

SSYN Siderone zethus Westwood, 1850
SSYN Zaretis isidora leopoldina Fruhstorfer, 1909
SSYN Siderone isidora naama Brévignon, 2009

SPEC Zaretis strigosus (Gmelin, [1790])

SSYN Siderone isidora strigosa Staudinger, 1887
SSYN Zaretis isidora russeus Fruhstorfer, 1909
SSYN Zaretis isidora vulpina Fruhstorfer, 1909
SSYN Zaretis isidora vulpecula Fruhstorfer, 1909
SSYN Siderone ellops mellita Brévignon, 2009

SPEC Zaretis itylus (Westwood, 1850)

SSYN Zaretis pseuditys Fruhstorfer, 1909

SPEC Zaretis ellops (Ménétriés, 1855)

SSYN Zaretis isidora anzuletta Fruhstorfer, 1909

SPEC Zaretis syene (Hewitson, 1856)
SPEC Zaretis pythagoras Willmott & J. Hall, 2004
SPEC Zaretis delassisei Choimet, 2009
SPEC Zaretis falcis Dias, Casagrande & Mielke, 2012
SPEC Zaretis hurin Dias, 2018
SPEC Zaretis elianahenrichae Dias, 2018
SPEC Zaretis mirandaenrichae Dias, 2018
SPEC Zaretis crawfordhilli Dias, 2018

GENU Phantos Dias, 2018

SPEC Phantos callidryas (R. Felder, 1869)
SPEC Phantos opalina (Godman & Salvin, 1884)

SSYN Siderone clara Staudinger, 1886

GENU Anaea Hübner, [1819]

GSYN Pyrrhanaea Röber, 1888

SPEC Anaea aidea (Guérin-Méneville, [1844])
SPEC Anaea andria Scudder, 1875
SPEC Anaea troglodyta (Fabricius, 1775)

SUBS Anaea troglodyta troglodyta (Fabricius, 1775)
SUBS Anaea troglodyta portia (Fabricius, 1775)
SUBS Anaea troglodyta minor A. Hall, 1936

SPEC Anaea astina (Fabricius, 1793)

SSYN Anaea troglodyta borinquenalis F. Johnson & W. Comstock, 1941

SPEC Anaea cubana (H. Druce, 1905)
SPEC Anaea floridalis F. Johnson & W. Comstock, 1941

GENU Fountainea Rydon, 1971

SPEC Fountainea centaurus (C. Felder & R. Felder, 1867)
SPEC Fountainea eurypyle (C. Felder & R. Felder, 1862)

SUBS Fountainea eurypyle eurypyle (C. Felder & R. Felder, 1862)
SUBS Fountainea eurypyle confusa (A. Hall, 1929)
SUBS Fountainea eurypyle glanzi (Rotger, Escalante & Coronado, 1965)

SPEC Fountainea glycerium (E. Doubleday, [1849])

SUBS Fountainea glycerium glycerium (E. Doubleday, [1849])
SUBS Fountainea glycerium comstocki (Witt, 1972)
SUBS Fountainea glycerium cratais (Hewitson, 1874)
SUBS Fountainea glycerium venezuelana (F. Johnson & W. Comstock, 1941)
SUBS Fountainea glycerium yucatanum (Witt, 1980)

SPEC Fountainea johnsoni (Avinoff & Shoumatoff, 1941)
SPEC Fountainea halice (Godart, [1824])

SUBS Fountainea halice halice (Godart, [1824])
SUBS Fountainea halice chrysophana (H. Bates, 1866)
SUBS Fountainea halice evelina (W. Comstock, 1961)
SUBS Fountainea halice fumata (A. Hall, 1935)
SUBS Fountainea halice martinezi (J. Maza & Díaz, 1978)
SUBS Fountainea halice maya (Witt, 1980)
SUBS Fountainea halice moretta (H. Druce, 1877)
SUBS Fountainea halice tehuana (A. Hall, 1917)

SPEC Fountainea nessus (Latreille, [1813])
SPEC Fountainea rayoensis (J. Maza & Díaz, 1978)
SPEC Fountainea nobilis (H. Bates, 1864)

SUBS Fountainea nobilis nobilis (H. Bates, 1864)
SUBS Fountainea nobilis caudata (Röber, 1916)
SUBS Fountainea nobilis pacifica (Vélez & Salazar, 1991)
SUBS Fountainea nobilis peralta (A. Hall, 1929)
SUBS Fountainea nobilis romeroi (Descimon, 1988)
SUBS Fountainea nobilis titan (C. Felder & R. Felder, 1867)

SPEC Fountainea ryphea (Cramer, 1775)

SUBS Fountainea ryphea ryphea (Cramer, 1775)
SUBS Fountainea ryphea ecuadoralis (F. Johnson & W. Comstock, 1941)
SUBS Fountainea ryphea phidile (Geyer, 1837)

SPEC Fountainea sosippus (Hopffer, 1874)

GENU Memphis Hübner, [1819]

GSYN Corycia Hübner, 1825
GSYN Cymatogramma Doubleday, 1849
GSYN Euschatzia Grote, 1898

SPEC Memphis acaudata (Röber, 1916)
SPEC Memphis acidalia (Hübner, [1819])

SUBS Memphis acidalia acidalia (Hübner, [1819])
SUBS Memphis acidalia memphis (C. Felder & R. Felder, 1867)
SUBS Memphis acidalia victoria (H. Druce, 1877)

SPEC Memphis alberta (H. Druce, 1876)
SPEC Memphis ambrosia (H. Druce, 1874)

SUBS Memphis ambrosia ambrosia (H. Druce, 1874)
SUBS Memphis ambrosia phoebe (H. Druce, 1877)

SPEC Memphis anassa (C. Felder & R. Felder, 1862)
SPEC Memphis anna (Staudinger, 1897)

SUBS Memphis anna anna (Staudinger, 1897)
SUBS Memphis anna elina (Staudinger, 1897)

SPEC Memphis appias (Hübner, [1825])
SPEC Memphis arginussa (Geyer, 1832)
SPEC Memphis onophis (C. Felder & R. Felder, 1861)
SPEC Memphis eubaena (Boisduval, 1870)
SPEC Memphis smalli Riley & Dias, 2021
SPEC Memphis artacaena (Hewitson, 1869)
SPEC Memphis aulica (Röber, 1916)
SPEC Memphis aureola (H. Bates, 1866)
SPEC Memphis basilia (Stoll, 1780)

SUBS Memphis basilia basilia (Stoll, 1780)
SUBS Memphis basilia bella (W. Comstock, 1961)
SUBS Memphis basilia drucei (Staudinger, 1887)

SPEC Memphis beatrix (H. Druce, 1874)
SPEC Memphis boliviana (H. Druce, 1877)
SPEC Memphis catinka (H. Druce, 1877)
SPEC Memphis cerealia (H. Druce, 1877)
SPEC Memphis cleomestra (Hewitson, 1869)

SUBS Memphis cleomestra cleomestra (Hewitson, 1869)
SUBS Memphis cleomestra ada (A. Butler, 1875)

SPEC Memphis cluvia (Hopffer, 1874)
SPEC Memphis dia (Godman & Salvin, 1884)

SUBS Memphis dia dia (Godman & Salvin, 1884)
SUBS Memphis dia divina (Röber, 1916)

SPEC Memphis falcata (Hopffer, 1874)
SPEC Memphis forreri (Godman & Salvin, 1884)
SPEC Memphis glauce (C. Felder & R. Felder, 1862)

SUBS Memphis glauce glauce (C. Felder & R. Felder, 1862)
SUBS Memphis glauce centralis (Röber, 1916)
SUBS Memphis glauce cicla (Möschler, 1877)
SUBS Memphis glauce felderi (Röber, 1916)

SPEC Memphis grandis (H. Druce, 1877)
SPEC Memphis hedemanni (R. Felder, 1869)
SPEC Memphis herbacea (A. Butler & H. Druce, 1872)
SPEC Memphis hirta (Weymer, 1907)
SPEC Memphis iphis (Latreille, [1813])
SPEC Memphis juliani Constantino, 1999
SPEC Memphis laertes (Cramer, 1775)
SPEC Memphis lankesteri (Hall, 1935)
SPEC Memphis laura (H. Druce, 1877)

SUBS Memphis laura laura (H. Druce, 1877)
SUBS Memphis laura balboa (A. Hall, 1927)
SUBS Memphis laura caucana (Joicey & Talbot, 1922)
SUBS Memphis laura rosae (Fassl, 1909)

SPEC Memphis lemnos (H. Druce, 1877)
SPEC Memphis leonida (Stoll, 1782)

SUBS Memphis leonida leonida (Stoll, 1782)
SUBS Memphis leonida editha (W. Comstock, 1961)

SPEC Memphis lineata (Salvin, 1869)
SPEC Memphis lorna (H. Druce, 1877)
SPEC Memphis lyceus (H. Druce, 1877)
SPEC Memphis maria Pyrcz & Neild, 1996
SPEC Memphis moeris (C. Felder & R. Felder, 1867)
SPEC Memphis montesino Pyrcz, 1995
SPEC Memphis mora (H. Druce, 1874)

SUBS Memphis mora mora (H. Druce, 1874)
SUBS Memphis mora annetta (W. Comstock, 1961)
SUBS Memphis mora montana (Röber, 1916)
SUBS Memphis mora orthesia (Godman & Salvin, 1884)

SPEC Memphis moruus (Fabricius, 1775)

SUBS Memphis moruus moruus (Fabricius, 1775)
SUBS Memphis moruus boisduvali (W. Comstock, 1961)
SUBS Memphis moruus leonila (W. Comstock, 1961)
SUBS Memphis moruus morpheus (Staudinger, [1886])
SUBS Memphis moruus phila (H. Druce, 1877)
SUBS Memphis moruus stheno (Prittwitz, 1865)

SPEC Memphis neidhoeferi (Rotger, Escalante & Coronado, 1965)
SPEC Memphis nenia (H. Druce, 1877)

SUBS Memphis nenia nenia (H. Druce, 1877)
SUBS Memphis nenia threnodion (Bargmann, 1929)

SPEC Memphis oenomais (Boisduval, 1870)
SPEC Memphis offa (H. Druce, 1877)

SUBS Memphis offa offa (H. Druce, 1877)
SUBS Memphis offa gudrun (Niepelt, 1924)

SPEC Memphis otrere (Hübner, [1825])
SPEC Memphis pasibula (E. Doubleday, [1849])
SPEC Memphis perenna (Godman & Salvin, 1884)

SUBS Memphis perenna perenna (Godman & Salvin, 1884)
SUBS Memphis perenna austrina (W. Comstock, 1961)
SUBS Memphis perenna lankesteri (A. Hall, 1935)

SPEC Memphis phantes (Hopffer, 1874)

SUBS Memphis phantes phantes (Hopffer, 1874)
SUBS Memphis phantes vicinia (Staudinger, 1887)

SPEC Memphis philumena (E. Doubleday, [1849])

SUBS Memphis philumena philumena (E. Doubleday, [1849])
SUBS Memphis philumena chaeronea (C. Felder & R. Felder, 1861)
SUBS Memphis philumena corita (Fruhstorfer, 1916)
SUBS Memphis philumena indigotica (Salvin, 1869)
SUBS Memphis philumena xenica (H. Bates, 1864)

SPEC Memphis pithyusa (R. Felder, 1869)

SSYN Anaea pithyusa morena Hall, 1935

SPEC Memphis polycarmes (Fabricius, 1775)
SPEC Memphis polyxo (H. Druce, 1874)
SPEC Memphis praxias (Hopffer, 1874)

SUBS Memphis praxias praxias (Hopffer, 1874)
SUBS Memphis praxias oblita (A. Hall, 1929)

SPEC Memphis schausiana (Godman & Salvin, 1894)
SPEC Memphis proserpina (Salvin, 1869)
SPEC Memphis elara (Godman & Salvin, 1897)
SPEC Memphis kingi (L. Miller & Nicolay, 1971)
SPEC Memphis pseudiphis (Staudinger, 1887)
SPEC Memphis salinasi Pyrcz, 1993
SPEC Memphis verticordia (Hübner, [1831])
SPEC Memphis echemus (E. Doubleday, [1849])

SUBS Memphis echemus echemus (E. Doubleday, [1849])
SUBS Memphis echemus bahamae (Witt, 1972)
SUBS Memphis echemus danieliana (Witt, 1972)

SPEC Memphis intermedia (Witt, 1972)

SUBS Memphis intermedia intermedia (Witt, 1972)
SUBS Memphis intermedia venus L. Miller, Simon & Harvey, 1992
SUBS Memphis intermedia mayaguanae L. Miller, Simon & Harvey, 1992

SPEC Memphis dominicana (Godman & Salvin, 1884)

SUBS Memphis dominicana dominicana (Godman & Salvin, 1884)
SUBS Memphis dominicana luciana (A. Hall, 1929)

SPEC Memphis viloriae Pyrcz & Neild, 1996
SPEC Memphis wellingi L. Miller & J. Miller, 1976
SPEC Memphis xenippa (A. Hall, 1935)
SPEC Memphis xenocles (Westwood, 1850)

SUBS Memphis xenocles xenocles (Westwood, 1850)
SUBS Memphis xenocles carolina (W. Comstock, 1961)
SUBS Memphis xenocles fisilis (A. Hall, 1935)
SUBS Memphis xenocles marginalis (A. Hall, 1935)

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