Harvey's classification of Nymphalidae

Niklas Wahlberg

Harvey, D. J. 1991. Higher classification of the Nymphalidae, Appendix B. - In: Nijhout, H. F. (ed) The Development and Evolution of Butterfly Wing Patterns. Smithsonian Institution Press, pp. 255-273.

This classification was the standard until recent molecular work made some crucial changes.

NYMPHALIDAE

I. HELICONIINAE
A. Pardopsini
Pardopsis
B. Acraeini
Acraea
Actinote
Cethosia

C. Heliconiini
1. Heliconiiti
Philaethria
Podotricha
Dione
Agraulis
Dryadula
Dryas
Eueides
Neruda
Laparus
Heliconius
2. Argynniti
Argyreus
Damora
Childrena
Pandoriana
Argynnis
Fabriciana
Speyeria
Mesoacidalia
Brenthis
Issoria
Argyronome
Nephargynnis
3. Boloriiti
Boloria
Clossiana
Proclossiana
Subtribe uncertain
Vindula
Cirrochroa
Paduca
Terinos
Lachnoptera
Smerina
Cupha
Vagrans
Phalanta
Euptoieta
Yramea
Kuekenthaliella
Prokuekenthaliella
II. NYMPHALINAE
A. Nymphalini
Vanessa
Cynthia
Bassaris
Aglais
Inachis
Nymphalis
Polygonia
Kaniska
Antanartia
Hypanartia
Symbrenthia
Mynes
Araschnia
B. Kallimini
Anartia
Metamorpha
Siproeta
Napeocles
Precis
Junonia
Hypolimnas
Salamis
Yoma
Kallima
Kallimoides
Doleschallia
Amnosia
Vanessula
Catacroptera
Rhinopalpa
C. Melitaeini
1. Euphydryiti
Euphydryas
Hypodryas
Occidryas
Eurodryas
2. Melitaeiti
Mellicta
Melitaea
Poladryas
Didymaeformia
Chlosyne
Thessalia
Texola
Dymasia
Microtia
Gnathotriche
Gnathotrusia
Higginsius
Antillea
3. Phycioditi
Phyciodes
Phystis
Anthanassa
Dagon
Telenassa
Ortilia
Tisonia
Tegosa
Eresia
Castilia
Janatella
Mazia
Subtribe uncertain
Atlantea
III. LIMENITIDINAE
A. Coloburini
Historis
Coea
Pycina
Baeotus
Smyrna
Colobura
Tigridia
B. Biblidini
1. Bibliditi
Biblis
2. Euryteliti
Mesoxantha
Ariadne
Eurytela
Neptidopsis
Laringa
Byblia
Mestra
Archimestra
Vila
3. Epicaliiti
Myscelia
Catonephele
Nessaea
Cybdelis
Eunica
Libythina
Sallya
4. Ageroniiti
Hamadryas
Ectima
Panacea
Batesia
5. Epiphiliti
Asterope
Pyrrhogyra
Temenis
Epiphile
Lucinia
Bolboneura
Nica
Peria
6. Dynaminiti
Dynamine
7. Catagrammiti
Diatheria
Perisama
Antigonis
Cyclogramma
Haematera
Paulogramma
Callicore
Catacore
C. Limenitidini
1. Limenitiditi
Limenitis
Parasarpa
Moduza
Sumalia
Auzakia
Adelpha
Basilarchia
Ladoga
Athyma
Pandita
Parathyma
Tacoraea
Cymothoe
Harma
Pseudacraea
Pseudoneptis
2. Neptiti
Pantoporia
Lasippa
Neptis
Phaedyma
Aldania
3. Partheniti
Parthenos
Lebadea
4. Euthaliiti
Tanaecia
Euthalia
Neurosigma
Abrota
Dophla
Lexias
Bassarona
Catuna
Hamanumida
Aterica
Cynandra
Euryphura
Pseudargynnis
Diestogyna
Euphaedra
Euriphene
Bebearia
D. Cyrestidini
Marpesia
Cyrestis
Chersonesia
Dichorragia
Incertae sedis
Pseudergolis
Stibochiona
IV. CHARAXINAE
A. Charaxini
Polyura
Murwareda
Charaxes
Haridra
Zingha
Stonehamia
Eriboea
B. Euxanthini
Euxanthe
Godartia
Hypomelaena
C. Pallini
Palla
D. Prothoini
Agatasa
Prothoe
E. Preponini
Anaeomorpha
Noreppa
Archaeoprepona
Prepona
Agrias
F. Anaeini
1. Zaretiditi
Coenophlebia
Zaretis
Siderone
2. Anaeiti
Hypna
Anaea
Polygrapha
Consul
Cymatogramma
Fountainea
Memphis
V. APATURINAE
Apatura
Apaturina
Apaturopsis
Asterocampa
Chitoria
Dilipa
Doxocopa
Euapatura
Eulaceura
Euripus
Helcyra
Herona
Hestina
Hestinalis
Mimathyma
Rohana
Sasakia
Sephisa
Timelaea
Thaleropis
VI. MORPHINAE
A. Morphini
1. Antirrheiti
Caerois
Antirrhea
2. Morphiti
Morpho
B. Amathusiini
Faunis
Aemona
Xanthotaenia
Taenaris
Morphotenaris
Stichophthalma
Amathusia
Amathuxidia
Zeuxidia
Thaumantis
Thauria
Discophora
Enispe
VII. BRASSOLINAE
Brassolis
Orobrassolis
Catoblepia
Dynastor
Opoptera
Mimoblepia
Opsiphanes
Mielkella
Penetes
Selenophanes
Caligo
Eryphanis
Dasyophthalma
Narope
Aponarope
VIII. SATYRINAE (contains several hundred genera and thus the classification goes to subtribal level only)
A. Haeterini
B. Biini
1. Biiti
2. Melanititi

C. Elymniini
1. Lethiti
2. Zetheriti
3. Elymniiti
4. Mycalesiti
D. Eritini
E. Ragadiini
F. Satyrini

1. Hypocystiti
2. Ypthimiti
3. Euptychiiti
4. Coenonymphiti
5. Manioliti
6. Erebiiti
7. Diriti
8. Pronophiliti
9. Satyriti
10. Melanargiiti
IX. CALINAGINAE
Calinaga
X. DANAINAE
A. Danaini
1. Amauriti
Parantica
Ideopsis
Amauris
2. Danaiti
Tirumala
Danaus
B. Euploeini
1. Euploeiti
Euploea
Idea
Protoploea
2. Ituniti
Lycorea
Anetia
XI. TELLERVINAE
Tellervo
XII. ITHOMINAE
A. Tithoreini
Elzunia
Tithorea
B. (New tribe)
Aeria
C. (New tribe)
Roswellia
Athesis
Patricia
D. Methonini
Methona
E. (New tribe)
Placidula
F. Melinaeini
Athyrtis
Olyras
Eutresis
Melinaea
G. Mechanitini
Paititia
Thyridia
Sais
Scada
Forbestra
Mechanitis
H. Oleriini
Hyposcada
(New genus)
Oleria
(New genus)
I. Napeogenini
Epityches
Rhodussa
Aremfoxia
Napeogenes
Garsauritis
Hyaliris
Hypothyris
J. Ithomiini
Miraleria
Pagyris
Ithomia
K. Dircennini
Callithomia
Velamysta
(New genus)
Dircenna
Hyalenna
L. (New tribe)
Ceratinia
Ceratiscada
Prittwitzia
Episcada
Pteronymia
M. Godyridini
Godyris
Dygoris
Pseudoscada
Hypomenitis
Greta
Hypoleria
Mcclungia
Veladyris
Heterosais
XIII. LIBYTHEINAE
Libythea
Libytheana