Pronophilina

Niklas Wahlberg

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Updated 2023-11-03. Based on

The Butterflies of America

Mahecha-J., O. et al 2021: Solving the cryptic diversity of the genus Manerebia Staudinger in northern Peru: description of new species and considerations on the biogeographical role of the Huancabamba Deflection (Nymphalidae: Satyrinae: Pronophilina). Zootaxa 5072: 201–237 DOI 10.11646/zootaxa.5072.3.1

Padrón, P.S., Pyrcz, T.W. & Willmott, K.R. 2021:A Contribution Towards Resolving the Systematics of the High-Altitude Tropical Andean Satyrine Genus Altopedaliodes Forster, 1964 (Lepidoptera, Nymphalinae: Satyrinae). Neotrop Entomol (2021). DOI 10.1007/s13744-021-00897-9

Pyrcz, T. W., Lorenc-Brudecka, J., Zubek, A., Boyer, P., Gabaldon, M. C. & Mavarez, J. 2017: Taxonomy, phylogeny and distribution of the genus Steromapedaliodes sensu novo in the Cordillera de Mérida, Venezuela (Lepidoptera: Nymphalidae: Satyrinae: Satyrini). Arthropod Systematics & Phylogeny 75: 195-243.

Pyrcz, T. W., Freitas, A. V. L., Boyer, P., Dias, F. M. S., Dolibaina, D. R., Barbosa, E. P., Magaldi, L. M., Mielke, O. H. H., Casagrande, M. M. & Lorenc-Brudecka, J. 2018:Uncovered diversity of a predominantly Andean butterfly clade in the Brazilian Atlantic forest: a revision of the genus Praepedaliodes Forster (Lepidoptera: Nymphalidae, Satyrinae, Satyrini). Neotropical Entomology 47: 211-255. DOI 10.1007/s13744-017-0543-x

Pyrcz, T. W., Cerdeña Gutierrez, J. & Florczyk, K. 2019: Two remarkable new species of Argyrophorus Blanchard from the Peruvian high Andes (Lepidoptera, Nymphalidae, Satyrinae). Zootaxa 4656: 375-383. DOI 10.11646/zootaxa.4656.2.10

Pyrcz, T.W., Stachowicz, I., Zubek, A. et al. 2021: A new species of butterfly of the genus Protopedaliodes from the isolated Tramen Tepui in the Guyana Shield (Lepidoptera, Nymphalidae, Satyrinae). Neotrop Entomol. DOI 10.1007/s13744-020-00830-6

Pyrcz, T.W., Lachowska-Cierlik, D., Willmott, K.R., Mrozek, A., Oscar Mahecha-Jiménez, , Fåhraeus, C., Boyer, P., Martín, S. & Espeland, M. 2022: A new genus in the diverse Andean Pedaliodes complex uncovered using target enrichment (Lepidoptera, Nymphalidae). Systematic Entomology, DOI 10.1111/syen.12568

Tomasz W Pyrcz, Keith R Willmott, Dorota Lachowska-Cierlik, Artur Mrozek, Klaudia Florczyk, Oscar Mahecha-J, Marianne Espeland 2023: Phylogeny and systematics of the “Pronophila clade,” with 2 new genera to resolve the formerly polyphyletic genus Pseudomaniola (Lepidoptera: Nymphalidae: Satyrinae), Insect Systematics and Diversity 7, Issue 5, 4, DOI 10.1093/isd/ixad020

Zhang, J., Cong, Q., Shen, J., Opler, P. A. and Grishin, N. V. 2021: Genomics-guided refinement of butterfly taxonomy. The Taxonomic Report 9-3.

SUBT Pronophilina

GENU Pronophila E. Doubleday, [1849]

SPEC Pronophila unifasciata Lathy, 1906

SUBS Pronophila unifasciata unifasciata Lathy, 1906
SUBS Pronophila unifasciata deverra Thieme, 1907
SUBS Pronophila unifasciata brennus Thieme, 1907
SUBS Pronophila unifasciata bogotensis Jurriaanse, 1926
SUBS Pronophila unifasciata australis Hayward, 1949
SUBS Pronophila unifasciata donachui Adams & Bernard, 1977
SUBS Pronophila unifasciata nanegalito Pyrcz, 2000
SUBS Pronophila unifasciata deceptoria Pyrcz, 2005
SUBS Pronophila unifasciata eugeni Pyrcz, 2005
SUBS Pronophila unifasciata kennethi Pyrcz, 2005
SUBS Pronophila unifasciata antioquiana Pyrcz, 2005

SPEC Pronophila juliani Adams & Bernard, 1977
SPEC Pronophila timanthes Salvin, 1871
SPEC Pronophila obscura Butler, 1868

SUBS Pronophila obscura obscura Butler, 1868
SUBS Pronophila obscura cuchillaensis Viloria & Camacho, 1999

SPEC Pronophila epidipnis Thieme, 1907

SUBS Pronophila epidipnis epidipnis Thieme, 1907
SUBS Pronophila epidipnis orchewitsoni Adams & Bernard, 1979
SUBS Pronophila epidipnis lathyi Pyrcz, 2000
SUBS Pronophila epidipnis orcidipnis Pyrcz & Viloria, 2007
SUBS Pronophila epidipnis perplexa Pyrcz, 2004

SPEC Pronophila tremocrata Pyrcz, 2004
SPEC Pronophila cordillera Westwood, 1851

SUBS Pronophila cordillera cordillera Westwood, 1851
SUBS Pronophila cordillera variabilis Butler, 1873
SUBS Pronophila cordillera colocasia Thieme, 1907

SPEC Pronophila orcus (Latreille, [1813])

SUBS Pronophila orcus orcus (Latreille, [1813])
SUBS Pronophila orcus locuples Weymer, 1912
SUBS Pronophila orcus nepete Thieme, 1907

SPEC Pronophila thelebe E. Doubleday, [1849]
SPEC Pronophila isobelae Pyrcz, 2000

SUBS Pronophila isobelae isobelae Pyrcz, 2000
SUBS Pronophila isobelae benevola Pyrcz, 2000

SPEC Pronophila attali Pyrcz, 2000
SPEC Pronophila rosenbergi Lathy, 1906

SUBS Pronophila rosenbergi rosenbergi Lathy, 1906
SUBS Pronophila rosenbergi puyango Pyrcz, 2000

SPEC Pronophila bernardi Pyrcz, 2004
SPEC Pronophila intercidona Thieme, 1907

SUBS Pronophila intercidona intercidona Thieme, 1907
SUBS Pronophila intercidona thelebina Thieme, 1907

SPEC Pronophila margarita Pyrcz, 2000

GENU Corades E. Doubleday, [1849]

GSYN Panarche Thieme 1907

SPEC Corades enyo Hewitson, [1849]

SUBS Corades enyo enyo Hewitson, [1849]
SUBS Corades enyo almo Thieme, 1907

SPEC Corades pax Watkins, 1939
SPEC Corades chelonis Hewitson, 1863

SUBS Corades chelonis chelonis Hewitson, 1863
SUBS Corades chelonis rubeta Thieme, 1907
SUBS Corades chelonis rectilinea Pyrcz & Viloria, 2005

SPEC Corades lactefusa Thieme, 1907
SPEC Corades iduna Hewitson, 1850

SUBS Corades iduna iduna Hewitson, 1850
SUBS Corades iduna procellaria Thieme, 1907
SUBS Corades iduna marginalis Butler, 1873
SUBS Corades iduna peruviana Butler, 1873
SUBS Corades iduna gradosi Lamas, 2003

SPEC Corades pannonia Hewitson, 1850

SUBS Corades pannonia pannonia Hewitson, 1850
SUBS Corades pannonia ploas Thieme, 1907
SUBS Corades pannonia condorita Lamas, [1997]
SUBS Corades pannonia albomaculata Staudinger, [1887]

SPEC Corades argentata Butler, 1868
SPEC Corades sareba Hewitson, 1850

SUBS Corades sareba sareba Hewitson, 1850
SUBS Corades sareba fusciplaga Butler, 1873

SPEC Corades cybele Butler, 1866

SUBS Corades cybele cybele Butler, 1866
SUBS Corades cybele semiplena Thieme, 1907

SPEC Corades medeba Hewitson, 1850

SUBS Corades medeba medeba Hewitson, 1850
SUBS Corades medeba columbina Staudinger, 1894
SUBS Corades medeba pittieri Pyrcz, 2010

SPEC Corades melania Staudinger, 1894

SUBS Corades melania melania Staudinger, 1894
SUBS Corades melania carlos Pyrcz, 2006

SPEC Corades cistene Hewitson, 1863

SUBS Corades cistene cistene Hewitson, 1863
SUBS Corades cistene generosa Thieme, 1907

SPEC Corades dymantis Thieme, 1907
SPEC Corades chirone Hewitson, 1863
SPEC Corades tripunctata Weymer, 1890

SUBS Corades tripunctata tripunctata Weymer, 1890
SUBS Corades tripunctata necrufa Pyrcz, 2004
SUBS Corades tripunctata malambo Pyrcz, 2006

SPEC Corades ulema Hewitson, 1850

SUBS Corades ulema ulema Hewitson, 1850
SUBS Corades ulema mirianae Lamas, [1997]

SPEC Corades anfortas (Thieme, 1907)

SUBS Corades anfortas anfortas (Thieme, 1907)
SUBS Corades anfortas celestini Pyrcz, 2005

SPEC Corades tricordatus Hewitson, 1874
SPEC Corades callipolis Hewitson, 1874
SPEC Corades liliaceus Pyrcz, 2005

GENU Daedalma Hewitson, 1858

SPEC Daedalma dinias Hewitson, 1858

SUBS Daedalma dinias dinias Hewitson, 1858
SUBS Daedalma dinias salmoni Pyrcz, 2011
SUBS Daedalma dinias emma Pyrcz & Greeney, 2011
SUBS Daedalma dinias leticia Pyrcz, 2011
SUBS Daedalma dinias romeliana Pyrcz & Prieto, 2011
SUBS Daedalma dinias granadillas Pyrcz & Willmott, 2011
SUBS Daedalma dinias radfordi Pyrcz & Willmott, 2011

SPEC Daedalma boliviana Staudinger, 1897

SUBS Daedalma boliviana boliviana Staudinger, 1897
SUBS Daedalma boliviana peruviana Pyrcz, 2004

SPEC Daedalma fraudata Pyrcz, 2004

SUBS Daedalma fraudata fraudata Pyrcz, 2004
SUBS Daedalma fraudata rubroreducta Pyrcz & Willmott, 2011

SPEC Daedalma drusilla Hewitson, 1858

SUBS Daedalma drusilla drusilla Hewitson, 1858
SUBS Daedalma drusilla tinta Pyrcz, 2011
SUBS Daedalma drusilla krugeriana Pyrcz, 2011

SPEC Daedalma inconspicua Butler, 1866

SUBS Daedalma inconspicua inconspicua Butler, 1866
SUBS Daedalma inconspicua blancae Pyrcz, 2011
SUBS Daedalma inconspicua cuencana Pyrcz, 2011
SUBS Daedalma inconspicua onorei Pyrcz, 2011
SUBS Daedalma inconspicua orientalis Pyrcz, 2011
SUBS Daedalma inconspicua variegata Pyrcz, 2011
SUBS Daedalma inconspicua palacio Dognin, 1891
SUBS Daedalma inconspicua petitia Pyrcz, 2011
SUBS Daedalma inconspicua tapichalaca Pyrcz & Willmott, 2011

SPEC Daedalma vertex Pyrcz, 2004

SUBS Daedalma vertex vertex Pyrcz, 2004
SUBS Daedalma vertex milleri Pyrcz, 2011

SPEC Daedalma parvomaculata E. Krüger, 1924
SPEC Daedalma adamsi D'Abrera, 1988
SPEC Daedalma eliza Pyrcz & Willmott, 2011
SPEC Daedalma dognini Pyrcz, 2011

SUBS Daedalma dognini dognini Pyrcz, 2011
SUBS Daedalma dognini mirianae Pyrcz, 2011

GENU Thiemeia Weymer, 1912

SPEC Thiemeia phoronea (E. Doubleday, [1849])

SUBS Thiemeia phoronea phoronea (E. Doubleday, [1849])
SUBS Thiemeia phoronea obscurata E. Krüger, 1924
SUBS Thiemeia phoronea ortruda (Thieme, 1907)

GENU Junea Hemming, 1964

GSYN Polymastus Thieme 1907

SPEC Junea dorinda (C. Felder & R. Felder, 1862)

SUBS Junea dorinda dorinda (C. Felder & R. Felder, 1862)
SUBS Junea dorinda quasinegra Pyrcz, 2004
SUBS Junea dorinda whitelyi (H. Druce, 1876)

SPEC Junea doraete (Hewitson, 1858)

SUBS Junea doraete doraete (Hewitson, 1858)
SUBS Junea doraete antissa (Thieme, 1907)
SUBS Junea doraete gideon (Thieme, 1907)
SUBS Junea doraete enipeus (Thieme, 1907)

GENU Oxeoschistus Butler, 1867

GSYN Dioriste Thieme 1907

SPEC Oxeoschistus protogenia (Hewitson, 1862)
SPEC Oxeoschistus pronax (Hewitson, 1859)
SPEC Oxeoschistus puerta (Westwood, 1851)

SUBS Oxeoschistus puerta submaculatus Butler & H. Druce, 1874
SUBS Oxeoschistus puerta pervius Thieme, 1907
SUBS Oxeoschistus puerta puerta (Westwood, 1851)
SUBS Oxeoschistus puerta magna Pyrcz & Viloria, 2007

SPEC Oxeoschistus simplex Butler, 1868

SUBS Oxeoschistus simplex simplex Butler, 1868
SUBS Oxeoschistus simplex fuscus R. Krüger, 1929
SUBS Oxeoschistus simplex isolda Thieme, 1907
SUBS Oxeoschistus simplex triplex Pyrcz & Rodriguez, MS
SUBS Oxeoschistus simplex duplex Godman, 1905
SUBS Oxeoschistus simplex garleppi Lamas, 2003

SPEC Oxeoschistus iphigenia Pyrcz, 2004
SPEC Oxeoschistus euriphyle Butler, 1872
SPEC Oxeoschistus hilara (H. Bates, 1865)
SPEC Oxeoschistus cothonides Grose-Smith, 1896
SPEC Oxeoschistus cothon Salvin, 1871
SPEC Oxeoschistus tauropolis (Westwood, [1850])
SPEC Oxeoschistus leucospilos Staudinger, 1876

SUBS Oxeoschistus leucospilos leucospilos Staudinger, 1876
SUBS Oxeoschistus leucospilos pugil (Thieme, 1907)

SPEC Oxeoschistus romeo Pyrcz & Fratello, 2005

GENU Mygona Thieme, 1907

SPEC Mygona irmina (E. Doubleday, [1849])
SPEC Mygona prochyta (Hewitson, 1862)
SPEC Mygona chyprota (Grose-Smith, 1900)
SPEC Mygona thammi (Staudinger, 1876)
SPEC Mygona poeania (Hewitson, 1870)

SUBS Mygona poeania poeania (Hewitson, 1870)
SUBS Mygona poeania magalyae Pyrcz, 2004

GENU Proboscis Thieme, 1907

SPEC Proboscis propylea (Hewitson, 1857)
SPEC Proboscis pomarancia Pyrcz, 2004

GENU Apexacuta Pyrcz, 2004

SPEC Apexacuta orsedice (Hewitson, 1878)

SUBS Apexacuta orsedice orsedice (Hewitson, 1878)
SUBS Apexacuta orsedice colombiana (E. Krüger, 1924)
SUBS Apexacuta orsedice violacea (Weymer, 1912)

SPEC Apexacuta improvisa Pyrcz, 2004
SPEC Apexacuta superior Pyrcz, 2004
SPEC Apexacuta astoreth (Thieme, 1907)

GENU Lasiophila C. Felder & R. Felder, 1859

SPEC Lasiophila cirta C. Felder & R. Felder, 1859

SUBS Lasiophila cirta cirta C. Felder & R. Felder, 1859
SUBS Lasiophila cirta atropurpurea Pyrcz, 2004

SPEC Lasiophila lucuma Pyrcz, 2011
SPEC Lasiophila piscina Thieme, 1903
SPEC Lasiophila luna Pyrcz, 2006
SPEC Lasiophila hewitsonia Butler, 1868

SUBS Lasiophila hewitsonia hewitsonia Butler, 1868
SUBS Lasiophila hewitsonia neda Thieme, 1907

SPEC Lasiophila zapatoza (Westwood, 1851)

SUBS Lasiophila zapatoza zapatoza (Westwood, 1851)
SUBS Lasiophila zapatoza meridae Adams & Bernard, 1981
SUBS Lasiophila zapatoza elusiva Pyrcz & Viloria, 2007
SUBS Lasiophila zapatoza manaurera Adams & Bernard, 1979
SUBS Lasiophila zapatoza sombra Thieme, 1907

SPEC Lasiophila semipartita Weymer, 1912
SPEC Lasiophila prosymna (Hewitson, 1857)

SUBS Lasiophila prosymna prosymna (Hewitson, 1857)
SUBS Lasiophila prosymna dirempta Thieme, 1907

SPEC Lasiophila parthyene (Hewitson, 1872)
SPEC Lasiophila ciris Thieme, 1907
SPEC Lasiophila circe C. Felder & R. Felder, 1859

SUBS Lasiophila circe circe C. Felder & R. Felder, 1859
SUBS Lasiophila circe farallonensis Pyrcz & Rodriguez, 2008
SUBS Lasiophila circe cnephas Thieme, 1907
SUBS Lasiophila circe arithmetica Pyrcz & Viloria, 2007
SUBS Lasiophila circe zarathustra Thieme, 1907

SPEC Lasiophila persepolis (Hewitson, 1872)
SPEC Lasiophila palades (Hewitson, 1872)

SUBS Lasiophila palades palades (Hewitson, 1872)
SUBS Lasiophila palades limes Pyrcz, 1999

SPEC Lasiophila phalaesia (Hewitson, 1868)

SUBS Lasiophila phalaesia phalaesia (Hewitson, 1868)
SUBS Lasiophila phalaesia alce Pyrcz, 1999
SUBS Lasiophila phalaesia confusa Staudinger, 1897
SUBS Lasiophila phalaesia kroli Pyrcz, 1999

SPEC Lasiophila regia Staudinger, 1897
SPEC Lasiophila orbifera Butler, 1868

SUBS Lasiophila orbifera orbifera Butler, 1868
SUBS Lasiophila orbifera munda Thieme, 1907
SUBS Lasiophila orbifera intercepta Thieme, 1907
SUBS Lasiophila orbifera pura Thieme, 1907

SPEC Lasiophila alkaios Tessmann, 1928

SUBS Lasiophila alkaios alkaios Tessmann, 1928
SUBS Lasiophila alkaios gita Smart, 1975

GENU Drucina Butler, 1872

SPEC Drucina championi Godman & Salvin, 1881
SPEC Drucina leonata Butler, 1872

GENU Druphila Pyrcz, 2004

SPEC Druphila venerata (A. Butler, 1873)

GENU Foetterleia Viloria, 2004

SPEC Foetterleia schreineri (Foetterle, 1902)

GENU Pseudomaniola Röber, 1889

GSYN Catargynnis Röber 1892

SPEC Pseudomaniola phaselis (Hewitson, 1862)
SPEC Pseudomaniola pholoe (Staudinger, 1887)
SPEC Pseudomaniola rogersi (Godman & Salvin, 1878)
SPEC Pseudomaniola macasica (Strand, 1912)
SPEC Pseudomaniola argyritis (Thieme, 1907)
SPEC Pseudomaniola gigas (Godman & Salvin, 1877)

GENU Boyeriana Pyrcz, Espeland & Willmott, 2023

SPEC Boyeriana mirabilis (A. Butler, 1873)
SPEC Boyeriana loxo (Dognin, 1891)
SPEC Boyeriana ilsa (Thieme, 1907)
SPEC Boyeriana extrema Pyrcz, 2004
SPEC Boyeriana gerlinda (Thieme, 1907)
SPEC Boyeriana mena (Grose-Smith, 1900)
SPEC Boyeriana clethra (Thieme, 1907)
SPEC Boyeriana helche (Thieme, 1907)

GENU Fahraeusia Pyrcz, 2023

SPEC Fahraeusia asuba (Thieme, 1907)

GENU Arhuaco Adams & Bernard, 1977

SPEC Arhuaco ica Adams & Bernard, 1977
SPEC Arhuaco dryadina (Schaus, 1913)

GENU Cheimas Thieme, 1907

SPEC Cheimas opalinus (Staudinger, 1897)

GENU Steroma Westwood, [1850]

GSYN Sierrasteroma Adams & Bernard 1977

SPEC Steroma bega Westwood, [1850]

SUBS Steroma bega bega Westwood, [1850]
SUBS Steroma bega andensis C. Felder & R. Felder, 1867

SPEC Steroma modesta Weymer, 1912
SPEC Steroma polyxo (Godman & Salvin, 1880)
SPEC Steroma superba Butler, 1868

GENU Steremnia Thieme, 1905

GSYN Pseudosteroma Weymer 1912

SPEC Steremnia lucillae Pyrcz, 2004
SPEC Steremnia monachella (Weymer, 1911)

SUBS Steremnia monachella monachella (Weymer, 1911)
SUBS Steremnia monachella adamsiella Pyrcz & Boyer, 2011

SPEC Steremnia pronophila (C. Felder & R. Felder, 1867)

SUBS Steremnia pronophila pronophila (C. Felder & R. Felder, 1867)
SUBS Steremnia pronophila fluminea Pyrcz & Viloria, 2007

SPEC Steremnia selva Adams, 1986
SPEC Steremnia umbracina (A. Butler, 1873)

SUBS Steremnia umbracina umbracina (A. Butler, 1873)
SUBS Steremnia umbracina misella Thieme, 1905

SPEC Steremnia agraulis (Weymer, 1912)

SUBS Steremnia agraulis agraulis (Weymer, 1912)
SUBS Steremnia agraulis agraulina Pyrcz, MS

SPEC Steremnia rugilas Thieme, 1905

GENU Eteona E. Doubleday, 1848

SPEC Eteona tisiphone (Boisduval, 1836)

GENU Eretris Thieme, 1905

SPEC Eretris oculata (C. Felder & R. Felder, 1867)
SPEC Eretris suzannae DeVries, 1980
SPEC Eretris lecromi Pyrcz, 1999
SPEC Eretris depresissima Pyrcz, 1999

SUBS Eretris depresissima depresissima Pyrcz, 1999
SUBS Eretris depresissima niambii Pyrcz, 1999

SPEC Eretris agata Pyrcz & Fratello, 2005
SPEC Eretris encycla (C. Felder & R. Felder, 1867)
SPEC Eretris neocycla Pyrcz & Viloria, 2009
SPEC Eretris calisto (C. Felder & R. Felder, 1867)

SUBS Eretris calisto calisto (C. Felder & R. Felder, 1867)
SUBS Eretris calisto kogui Adams & Bernard, 1977

SPEC Eretris ocellifera (C. Felder & R. Felder, 1867)
SPEC Eretris hulda (A. Butler & H. Druce, 1872)
SPEC Eretris maria (Schaus, 1920)
SPEC Eretris centralis E. Krüger, 1924
SPEC Eretris porphyria (C. Felder & R. Felder, 1867)

SUBS Eretris porphyria porphyria (C. Felder & R. Felder, 1867)
SUBS Eretris porphyria decorata (C. Felder & R. Felder, 1867)
SUBS Eretris porphyria catargyrea (Staudinger, 1887)
SUBS Eretris porphyria perija Adams & Bernard, 1979
SUBS Eretris porphyria recta Pyrcz & Viloria, 2007
SUBS Eretris porphyria pseudoperija Pyrcz, 1999
SUBS Eretris porphyria transmaraniona Pyrcz, 2004

SPEC Eretris apuleja (C. Felder & R. Felder, 1867)

SUBS Eretris apuleja apuleja (C. Felder & R. Felder, 1867)
SUBS Eretris apuleja subrufescens (Grose-Smith & W. F. Kirby, 1895)
SUBS Eretris apuleja bogotana E. Krüger, 1924
SUBS Eretris apuleja ochrea Thieme, 1905
SUBS Eretris apuleja altamira Adams & Bernard, 1979
SUBS Eretris apuleja remotissima Pyrcz & Viloria, 2007
SUBS Eretris apuleja socorrensis Pyrcz & Rodriguez, 2008

SPEC Eretris microrufescens Pyrcz & Wojtusiak, 2010
SPEC Eretris suprarufescens Pyrcz & Wojtusiak, 2010
SPEC Eretris apuleina (Pyrcz, 2004)
SPEC Eretris truncatina (Pyrcz, 2004)
SPEC Eretris subpunctata (Grose-Smith & W. F. Kirby, 1895)
SPEC Eretris mendoza (Pyrcz, 2004)
SPEC Eretris julieta Pyrcz & Gareca, 2010

GENU Manerebia Staudinger, 1897

GSYN Penrosada Brown 1944
GSYN Posteuptychia Forster 1964

SPEC Manerebia leaena (Hewitson, 1861)

SUBS Manerebia leaena leaena (Hewitson, 1861)
SUBS Manerebia leaena lanassa (C. Felder & R. Felder, 1867)
SUBS Manerebia leaena gonzalezi Pyrcz & Viloria, 2006

SPEC Manerebia inderena (Adams, 1986)

SUBS Manerebia inderena inderena (Adams, 1986)
SUBS Manerebia inderena antioquiana Pyrcz & Willmott, 2006
SUBS Manerebia inderena fina Pyrcz & Willmott, 2006
SUBS Manerebia inderena similis Pyrcz & Willmott, 2006
SUBS Manerebia inderena clara Pyrcz & Willmott, 2006
SUBS Manerebia inderena leaeniva Pyrcz & Willmott, 2006
SUBS Manerebia inderena mirena Pyrcz & Willmott, 2006

SPEC Manerebia prattorum Pyrcz & Willmott, 2006

SUBS Manerebia prattorum prattorum Pyrcz & Willmott, 2006
SUBS Manerebia prattorum udima Pyrcz & Boyer, 2021

SPEC Manerebia golondrina Pyrcz & Willmott, 2006
SPEC Manerebia trimaculata (Hewitson, 1870)
SPEC Manerebia undulata Pyrcz & J. Hall, 2006

SUBS Manerebia undulata undulata Pyrcz & J. Hall, 2006
SUBS Manerebia undulata milaena Pyrcz & Willmott, 2006

SPEC Manerebia interrupta (F. Brown, 1944)
SPEC Manerebia rufanalis Pyrcz & J. Hall, 2006

SUBS Manerebia rufanalis rufanalis Pyrcz & J. Hall, 2006
SUBS Manerebia rufanalis fernandina Pyrcz & Willmott, 2006

SPEC Manerebia quinterae (Adams & Bernard, 1979)
SPEC Manerebia navarrae (Adams & Bernard, 1979)
SPEC Manerebia satura (Weymer, 1911)

SUBS Manerebia satura satura (Weymer, 1911)
SUBS Manerebia satura keithi (Dyar, 1913)
SUBS Manerebia satura pauperata Pyrcz & Willmott, 2006
SUBS Manerebia satura lamasi Pyrcz & Willmott, 2006

SPEC Manerebia germaniae Pyrcz & J. Hall, 2006

SUBS Manerebia germaniae germaniae Pyrcz & J. Hall, 2006
SUBS Manerebia germaniae vitalei Pyrcz & Willmott, 2006

SPEC Manerebia mammuthus Pyrcz & Willmott, 2006
SPEC Manerebia franciscae (Adams & Bernard, 1981)

SUBS Manerebia franciscae franciscae (Adams & Bernard, 1981)
SUBS Manerebia franciscae rodriguezi Pyrcz & Willmott, 2006

SPEC Manerebia nevadensis E. Krüger, 1925
SPEC Manerebia mycalesoides (C. Felder & R. Felder, 1867)
SPEC Manerebia magnifica Pyrcz & Willmott, 2006
SPEC Manerebia levana (Godman, 1905)

SUBS Manerebia levana levana (Godman, 1905)
SUBS Manerebia levana pervaga Pyrcz & Viloria, 2007

SPEC Manerebia apiculata (C. Felder & R. Felder, 1867)
SPEC Manerebia ignilineata (Dognin, 1896)

SUBS Manerebia ignilineata ignilineata (Dognin, 1896)
SUBS Manerebia ignilineata neglecta (F. Brown, 1944)
SUBS Manerebia ignilineata jalca Pyrcz, 2004

SPEC Manerebia seducta Pyrcz & Willmott, 2006
SPEC Manerebia diffusa Pyrcz, 2004
SPEC Manerebia zoippus (H. Druce, 1876)
SPEC Manerebia benigni Pyrcz, 2004

SUBS Manerebia benigni benigni Pyrcz, 2004
SUBS Manerebia benigni tessmanni Pyrcz, 2004

SPEC Manerebia trirufa Pyrcz, 2004
SPEC Manerebia haywardi Pyrcz, 2004
SPEC Manerebia rubescens (A. Butler, 1873)
SPEC Manerebia cyclopella Staudinger, 1897
SPEC Manerebia cyclops Staudinger, 1897
SPEC Manerebia cyclopina Staudinger, 1897

SUBS Manerebia cyclopina cyclopina Staudinger, 1897
SUBS Manerebia cyclopina staudingeri Forster, 1964

SPEC Manerebia monops Hayward, 1949
SPEC Manerebia insulsa (Hewitson, 1874)
SPEC Manerebia typhlops Staudinger, 1897
SPEC Manerebia pluviosa Pyrcz & Viloria, 2007
SPEC Manerebia reducta (F. Brown, 1944)
SPEC Manerebia inducta Pyrcz & Willmott, 2021
SPEC Manerebia ronda Pyrcz & Boyer, 2021

SUBS Manerebia ronda ronda Pyrcz & Boyer, 2021
SUBS Manerebia ronda amplia Pyrcz & Boyer, 2021

SPEC Manerebia punku Pyrcz & Farfán, 2021
SPEC Manerebia huamanii Cerdeña & Pyrcz, 2021
SPEC Manerebia placida Pyrcz & Willmott, 2021
SPEC Manerebia granatus Willmott, Radford & Pyrcz, 2021

GENU Diaphanos Adams & Bernard, 1981

SPEC Diaphanos huberi Adams & Bernard, 1981
SPEC Diaphanos curvignathos Viloria, 1994
SPEC Diaphanos fuscus Viloria, 1994

GENU Idioneurula Strand, 1932

GSYN Idioneura Felder & Felder 1867

SPEC Idioneurula erebioides (C. Felder & R. Felder, 1867)

SUBS Idioneurula erebioides erebioides (C. Felder & R. Felder, 1867)
SUBS Idioneurula erebioides socorroi Villalobos-Moreno & Salazar, 2013

SPEC Idioneurula donegani (Huertas & Arias, 2008)
SPEC Idioneurula eremita Pyrcz & Viloria, 2007
SPEC Idioneurula jacquelinae (Pyrcz, 1995)

GENU Lymanopoda Westwood, 1851

GSYN Sarromia Westwood 1851
GSYN Zabirnia Hewitson 1877
GSYN Trophonina Röber 1889
GSYN Sabatoga Staudinger 1897

SPEC Lymanopoda panacea (Hewitson, 1869)

SUBS Lymanopoda panacea panacea (Hewitson, 1869)
SUBS Lymanopoda panacea gortyna (Weymer, 1890)
SUBS Lymanopoda panacea ocellifera Butler, 1873
SUBS Lymanopoda panacea venusia Hopffer, 1874

SPEC Lymanopoda affineola Weymer, 1911
SPEC Lymanopoda albomaculata Hewitson, 1861
SPEC Lymanopoda albocincta Hewitson, 1861

SUBS Lymanopoda albocincta albocincta Hewitson, 1861
SUBS Lymanopoda albocincta issacha Butler, 1870
SUBS Lymanopoda albocincta intermedia Pyrcz, 2004

SPEC Lymanopoda florenciaensis Salazar, Henao & Vargas, 2004
SPEC Lymanopoda apulia Hopffer, 1874
SPEC Lymanopoda caucana Weymer, 1911

SUBS Lymanopoda caucana caucana Weymer, 1911
SUBS Lymanopoda caucana orientalis Viloria & Camacho, 1999

SPEC Lymanopoda caeruleata Godman & Salvin, 1880
SPEC Lymanopoda euopis Godman & Salvin, 1878
SPEC Lymanopoda cinna Godman & Salvin, 1889
SPEC Lymanopoda acraeida Butler, 1868

SUBS Lymanopoda acraeida acraeida Butler, 1868
SUBS Lymanopoda acraeida zigomala (Hewitson, 1877)
SUBS Lymanopoda acraeida chavezi Pyrcz, 2004
SUBS Lymanopoda acraeida malia Godman, 1905

SPEC Lymanopoda venosa Butler, 1868
SPEC Lymanopoda obsoleta (Westwood, 1851)
SPEC Lymanopoda altis Weymer, 1890
SPEC Lymanopoda maletera Adams & Bernard, 1979
SPEC Lymanopoda dietzi Adams & Bernard, 1981

SUBS Lymanopoda dietzi dietzi Adams & Bernard, 1981
SUBS Lymanopoda dietzi neildi Pyrcz, 2010
SUBS Lymanopoda dietzi vetula Viloria & Pyrcz, 2010
SUBS Lymanopoda dietzi oliva Pyrcz & Boyer, 2010
SUBS Lymanopoda dietzi rosanna Viloria & Pyrcz, 2010
SUBS Lymanopoda dietzi argentata Pyrcz, 2010

SPEC Lymanopoda confusa F. Brown, 1943
SPEC Lymanopoda lecromi Pyrcz & Viloria, 2007
SPEC Lymanopoda confusiana Pyrcz & Boyer, 2010
SPEC Lymanopoda magna Pyrcz, 2004
SPEC Lymanopoda caudalis Rosenberg & Talbot, 1914

SUBS Lymanopoda caudalis caudalis Rosenberg & Talbot, 1914
SUBS Lymanopoda caudalis raymondae Pyrcz & Boyer, 2010

SPEC Lymanopoda balintiana Pyrcz & Boyer, 2010
SPEC Lymanopoda inaudita Pyrcz, 2010
SPEC Lymanopoda necalba Pyrcz & Boyer, 2010
SPEC Lymanopoda marianna Staudinger, 1897

SUBS Lymanopoda marianna marianna Staudinger, 1897
SUBS Lymanopoda marianna veronicae Pyrcz, 2010
SUBS Lymanopoda marianna giselae Pyrcz, 2010
SUBS Lymanopoda marianna ioanae Pyrcz, 2010
SUBS Lymanopoda marianna jacquelinae Pyrcz, 2010
SUBS Lymanopoda marianna valentinae Pyrcz, 2010

SPEC Lymanopoda labda Hewitson, 1861

SUBS Lymanopoda labda labda Hewitson, 1861
SUBS Lymanopoda labda moritzi Pyrcz & Viloria, 2007
SUBS Lymanopoda labda lebbaea C. Felder & R. Felder, 1867
SUBS Lymanopoda labda bilinskii Pyrcz, 2011

SPEC Lymanopoda altaselva Adams & Bernard, 1979
SPEC Lymanopoda nadia Pyrcz, 1999
SPEC Lymanopoda ionius Westwood, 1851

SUBS Lymanopoda ionius ionius Westwood, 1851
SUBS Lymanopoda ionius excisa Weymer, 1911
SUBS Lymanopoda ionius browni Pyrcz, 1999
SUBS Lymanopoda ionius gargantua Pyrcz & Rodriguez, 2008

SPEC Lymanopoda lactea Hewitson, 1861
SPEC Lymanopoda schmidti Adams, 1986
SPEC Lymanopoda labineta Hewitson, 1870

SUBS Lymanopoda labineta labineta Hewitson, 1870
SUBS Lymanopoda labineta piniasi Pyrcz, 1999
SUBS Lymanopoda labineta decorata Seydel, 1924
SUBS Lymanopoda labineta wojtusiaki Pyrcz, 2011

SPEC Lymanopoda pieridina Röber, 1927

SUBS Lymanopoda pieridina pieridina Röber, 1927
SUBS Lymanopoda pieridina albicosta Pyrcz, 1999

SPEC Lymanopoda paisa Pyrcz & Rodriguez, 2006
SPEC Lymanopoda nivea Staudinger, 1887

SUBS Lymanopoda nivea nivea Staudinger, 1887
SUBS Lymanopoda nivea sororcula Thieme, 1904
SUBS Lymanopoda nivea bingo Pyrcz, 1999
SUBS Lymanopoda nivea bonita Pyrcz, 1999

SPEC Lymanopoda samius Westwood, 1851

SUBS Lymanopoda samius samius Westwood, 1851
SUBS Lymanopoda samius lineana Pyrcz & Viloria, 2007

SPEC Lymanopoda ferruginosa Butler, 1868
SPEC Lymanopoda prusia (Heimlich, 1973)
SPEC Lymanopoda rana Weymer, 1911

SUBS Lymanopoda rana rana Weymer, 1911
SUBS Lymanopoda rana marrana Pyrcz & Boyer, 2011

SPEC Lymanopoda hyagnis Weymer, 1911
SPEC Lymanopoda translucida Weymer, 1912

SUBS Lymanopoda translucida translucida Weymer, 1912
SUBS Lymanopoda translucida umbratilis Rosenberg & Talbot, 1914
SUBS Lymanopoda translucida leucotecta Rosenberg & Talbot, 1914

SPEC Lymanopoda hockingiana Pyrcz, 2011
SPEC Lymanopoda araneola Pyrcz, 2004
SPEC Lymanopoda galactea Staudinger, 1897
SPEC Lymanopoda shefteli Dyar, 1913

SUBS Lymanopoda shefteli shefteli Dyar, 1913
SUBS Lymanopoda shefteli siviae Fuchs, 1954
SUBS Lymanopoda shefteli broweri Pyrcz & Boyer, 2011

SPEC Lymanopoda mariposa Pyrcz & Boyer, 2011
SPEC Lymanopoda ingasayana Pyrcz, 2004
SPEC Lymanopoda dyari Pyrcz, 2004
SPEC Lymanopoda inde Pyrcz, 2004
SPEC Lymanopoda eubagioides Butler, 1873
SPEC Lymanopoda hazelana F. Brown, 1943

SUBS Lymanopoda hazelana hazelana F. Brown, 1943
SUBS Lymanopoda hazelana summa Pyrcz, 1999
SUBS Lymanopoda hazelana viridia Pyrcz, 2011

SPEC Lymanopoda melia Weymer, 1911

SUBS Lymanopoda melia melia Weymer, 1911
SUBS Lymanopoda melia alba E. Krüger, 1924

SPEC Lymanopoda pascua Pyrcz, 2005
SPEC Lymanopoda tolima Weymer, 1911
SPEC Lymanopoda huilana Weymer, 1890

SUBS Lymanopoda huilana huilana Weymer, 1890
SUBS Lymanopoda huilana cajamarca Adams, 1986
SUBS Lymanopoda huilana dominicae Pyrcz, MS

SPEC Lymanopoda caracara Pyrcz, Willmott & J. Hall, 1999

SUBS Lymanopoda caracara caracara Pyrcz, Willmott & J. Hall, 1999
SUBS Lymanopoda caracara ichu Pyrcz, Willmott & J. Hall, 1999

SPEC Lymanopoda viventieni (Apolinar, 1924)
SPEC Lymanopoda mirabilis Staudinger, 1897
SPEC Lymanopoda paramera Adams & Bernard, 1979
SPEC Lymanopoda nevada E. Krüger, 1924

SUBS Lymanopoda nevada nevada E. Krüger, 1924
SUBS Lymanopoda nevada nevadensis (A. Schultze, 1931)

SPEC Lymanopoda melendeza Adams, 1986

GENU Ianussiusa Pyrcz & Viloria, 2004

SPEC Ianussiusa maso (Godman, 1905)

SUBS Ianussiusa maso maso (Godman, 1905)
SUBS Ianussiusa maso santa Pyrcz & Viloria, 2007
SUBS Ianussiusa maso polyommatus (Röber, 1927)

GENU Dangond Adams & Bernard, 1979

SPEC Dangond dangondi Adams & Bernard, 1979

GENU Altopedaliodes Forster, 1964

SPEC Altopedaliodes reissi (Weymer, 1890)

SUBS Altopedaliodes reissi reissi (Weymer, 1890)
SUBS Altopedaliodes reissi flavomaculata (E. Krüger, 1924)
SUBS Altopedaliodes reissi salazari Le Crom, 1994
SUBS Altopedaliodes reissi papallacta Padrón, Pyrcz and Willmott, 2021
SUBS Altopedaliodes reissi dominica Pyrcz and Padrón, 2021

SPEC Altopedaliodes halli Pyrcz, 2004

SUBS Altopedaliodes halli halli Pyrcz, 2004
SUBS Altopedaliodes halli cagnoni Boyer, Pyrcz and Padrón, 2021

SPEC Altopedaliodes belmira Pyrcz & Rodriguez, 2004
SPEC Altopedaliodes scydmaena (Hayward, 1968)

SUBS Altopedaliodes scydmaena kruegeri Pyrcz, 1999

SPEC Altopedaliodes tena (Hewitson, 1869)
SPEC Altopedaliodes nucea Pyrcz & Viloria, 1999
SPEC Altopedaliodes pasicles (Hewitson, 1872)
SPEC Altopedaliodes perita (Hewitson, 1868)

SUBS Altopedaliodes perita perita (Hewitson, 1868)
SUBS Altopedaliodes perita sorda Pyrcz, 2004

SPEC Altopedaliodes zsolti Pyrcz & Viloria, 1999

SUBS Altopedaliodes zsolti zsolti Pyrcz & Viloria, 1999
SUBS Altopedaliodes zsolti citra Pyrcz, 2004

SPEC Altopedaliodes kurti Pyrcz & Viloria, 1999
SPEC Altopedaliodes llanganati Padrón, Pyrcz and Willmott, 2021
SPEC Altopedaliodes pilimbala Pyrcz and Boyer, 2021

GENU Punapedaliodes Forster, 1964

SPEC Punapedaliodes flavopunctata (Staudinger, 1894)

SUBS Punapedaliodes flavopunctata flavopunctata (Staudinger, 1894)
SUBS Punapedaliodes flavopunctata minima Pyrcz, 2004

GENU Antopedaliodes Forster, 1964

SPEC Antopedaliodes albopunctata (Weymer, 1890)
SPEC Antopedaliodes antonia (Staudinger, 1897)

SUBS Antopedaliodes antonia antonia (Staudinger, 1897)
SUBS Antopedaliodes antonia anina (Staudinger, 1897)
SUBS Antopedaliodes antonia quincedis (Thieme, 1905)

GENU Corderopedaliodes Forster, 1964

SPEC Corderopedaliodes corderoi (Dognin, 1893)

SUBS Corderopedaliodes corderoi corderoi (Dognin, 1893)
SUBS Corderopedaliodes corderoi exornata Pyrcz, 2004

SPEC Corderopedaliodes buda Pyrcz, 2013
SPEC Corderopedaliodes luzangelae Pyrcz & Warren-Gash, 2013
SPEC Corderopedaliodes pandates (Hewitson, 1874)

SUBS Corderopedaliodes pandates pandates (Hewitson, 1874)
SUBS Corderopedaliodes pandates olivieri Pyrcz, 2013

GENU Panyapedaliodes Forster, 1964

GSYN Muscopedaliodes Forster 1964

SPEC Panyapedaliodes drymaea (Hewitson, 1858)

SUBS Panyapedaliodes drymaea drymaea (Hewitson, 1858)
SUBS Panyapedaliodes drymaea atropulla Pyrcz, 2005

SPEC Panyapedaliodes jephtha (Thieme, 1905)
SPEC Panyapedaliodes mara (Thieme, 1905)
SPEC Panyapedaliodes monticola (Tessmann, 1928)
SPEC Panyapedaliodes muscosa (Thieme, 1905)

SUBS Panyapedaliodes muscosa muscosa (Thieme, 1905)
SUBS Panyapedaliodes muscosa sumaco Pyrcz & Rodriguez, 2005
SUBS Panyapedaliodes muscosa tambito Pyrcz & Rodriguez, 2005

SPEC Panyapedaliodes panyasis (Hewitson, 1862)
SPEC Panyapedaliodes stellata Pyrcz, 2004
SPEC Panyapedaliodes phila (Hewitson, 1862)

SUBS Panyapedaliodes phila phila (Hewitson, 1862)
SUBS Panyapedaliodes phila certa Pyrcz, 2004
SUBS Panyapedaliodes phila combeima (E. Krüger, 1924)
SUBS Panyapedaliodes phila philaenis (Thieme, 1905)
SUBS Panyapedaliodes phila pilla Lamas, 2003

SPEC Panyapedaliodes puma (Thieme, 1905)

SUBS Panyapedaliodes puma puma (Thieme, 1905)
SUBS Panyapedaliodes puma ernsti Pyrcz, 2006

SPEC Panyapedaliodes rahab (Thieme, 1905)
SPEC Panyapedaliodes silpa (Thieme, 1905)

SUBS Panyapedaliodes silpa silpa (Thieme, 1905)
SUBS Panyapedaliodes silpa tomentosa (Weymer, 1912)

SPEC Panyapedaliodes traceyannae Pyrcz & Viloria, 1999
SPEC Panyapedaliodes cocorna Pyrcz & Rodriguez, 2005

GENU Paramo Adams & Bernard, 1977

SPEC Paramo oculata (E. Krüger, 1924)

GENU Parapedaliodes Forster, 1964

SPEC Parapedaliodes parepa (Hewitson, 1862)

SUBS Parapedaliodes parepa parepa (Hewitson, 1862)
SUBS Parapedaliodes parepa milvia (Thieme, 1905)

GENU Pherepedaliodes Forster, 1964

SPEC Pherepedaliodes naevia (Thieme, 1905)
SPEC Pherepedaliodes nubilia Pyrcz & Viloria, 1999
SPEC Pherepedaliodes angela Pyrcz & Rodriguez, 2005
SPEC Pherepedaliodes pheretiades (Grose-Smith & W. F. Kirby, 1894)

GENU Physcopedaliodes Forster, 1964

SPEC Physcopedaliodes physcoa (Hewitson, 1862)

SUBS Physcopedaliodes physcoa physcoa (Hewitson, 1862)
SUBS Physcopedaliodes physcoa marulla (Thieme, 1905)
SUBS Physcopedaliodes physcoa micromaculata Forster, 1964

GENU Praepedaliodes Forster, 1964

SPEC Praepedaliodes amussis (Thieme, 1905)
SPEC Praepedaliodes granulata (A. Butler, 1868)
SPEC Praepedaliodes phanias (Hewitson, 1862)
SPEC Praepedaliodes exul (Thieme, 1905)
SPEC Praepedaliodes duartei Dias, Dolibaina & Pyrcz, 2018
SPEC Praepedaliodes zaccae Dolibaina, Dias & Pyrcz, 2018
SPEC Praepedaliodes francinii Freitas & Pyrcz, 2018
SPEC Praepedaliodes sequeirae Pyrcz, Dias & Dolbaina, 2018
SPEC Praepedaliodes landryi Pyrcz & Freitas, 2018
SPEC Praepedaliodes pawlaki Pyrcz & Boyer, 2018

GENU Praepronophila Forster, 1964

SPEC Praepronophila emma (Staudinger, 1897)
SPEC Praepronophila perperna (Hewitson, 1862)

SUBS Praepronophila perperna perperna (Hewitson, 1862)
SUBS Praepronophila perperna smalli Pyrcz & Viloria, 2007
SUBS Praepronophila perperna rodriguezi Pyrcz & Viloria, 2007

SPEC Praepronophila petronius (Grose-Smith, 1900)

SUBS Praepronophila petronius kerrianna (L. Miller, 1986)
SUBS Praepronophila petronius petronius (Grose-Smith, 1900)

GENU Steromapedaliodes Forster, 1964

GSYN Redonda Adams & Bernard, 1981

SPEC Steromapedaliodes albonotata (Godman, 1905)

SUBS Steromapedaliodes albonotata albonotata (Godman, 1905)
SUBS Steromapedaliodes albonotata thiemeiana Pyrcz, 2017
SUBS Steromapedaliodes albonotata sanchezi Viloria & Pyrcz, 2001
SUBS Steromapedaliodes albonotata schuberti Viloria & Pyrcz, 2001
SUBS Steromapedaliodes albonotata australis Pyrcz & Boyer, 2017
SUBS Steromapedaliodes albonotata pratense Pyrcz & Boyer, 2017
SUBS Steromapedaliodes albonotata josefinae Pyrcz & Boyer, 2017
SUBS Steromapedaliodes albonotata borealis Pyrcz & Boyer, 2017

SPEC Steromapedaliodes albarregas (Adams & Bernard, 1981)

SUBS Steromapedaliodes albarregas albarregas (Adams & Bernard, 1981)
SUBS Steromapedaliodes albarregas mauroi Pyrcz, 2017
SUBS Steromapedaliodes albarregas prietoi Pyrcz, 2017

SPEC Steromapedaliodes kahlua Pyrcz & Boyer, 2017
SPEC Steromapedaliodes mavarezi Pyrcz, 2017
SPEC Steromapedaliodes empetrus (Thieme, 1905)

SUBS Steromapedaliodes empetrus empetrus (Thieme, 1905)
SUBS Steromapedaliodes empetrus bolivari (Adams & Bernard, 1981)
SUBS Steromapedaliodes empetrus frailejona (Ferrer-Paris & Costa, 2015)

SPEC Steromapedaliodes chiquinquirana (Ferrer-Paris, 2015)

SUBS Steromapedaliodes chiquinquirana chiquinquirana (Ferrer-Paris, 2015)
SUBS Steromapedaliodes chiquinquirana lossadana (Ferrer-Paris, 2015)
SUBS Steromapedaliodes chiquinquirana castellana (Viloria & Camacho, 2015)

SPEC Steromapedaliodes leukasmena (Viloria & Camacho, 2015)
SPEC Steromapedaliodes bordoni (Viloria & Pyrcz, 2003)
SPEC Steromapedaliodes lathraia (Viloria & Camacho, 2015)

GENU Neopedaliodes Viloria, L. Miller & J. Miller, 2004

SPEC Neopedaliodes chingazaensis (Torres & Le Crom, 1997)
SPEC Neopedaliodes zipa (Adams, 1986)
SPEC Neopedaliodes nora (Adams, 1986)

SUBS Neopedaliodes nora nora (Adams, 1986)
SUBS Neopedaliodes nora stepheni Viloria, L. Miller & J. Miller, 2008

SPEC Neopedaliodes margaretha (Adams, 1986)

SUBS Neopedaliodes margaretha margaretha (Adams, 1986)
SUBS Neopedaliodes margaretha allyni Viloria, L. Miller & J. Miller, 2008

SPEC Neopedaliodes philotera (Hewitson, 1868)
SPEC Neopedaliodes lefebvreae Viloria, L. Miller & J. Miller, 2008
SPEC Neopedaliodes parrhoebia (Hewitson, 1872)

SUBS Neopedaliodes parrhoebia parrhoebia (Hewitson, 1872)
SUBS Neopedaliodes parrhoebia phintia (Hewitson, 1877)

SPEC Neopedaliodes entella (Thieme, 1905)
SPEC Neopedaliodes juba (Staudinger, 1887)

SUBS Neopedaliodes juba juba (Staudinger, 1887)
SUBS Neopedaliodes juba jubilosa Viloria, L. Miller & J. Miller, 2008

SPEC Neopedaliodes michaeli Viloria, L. Miller & J. Miller, 2008

SUBS Neopedaliodes michaeli michaeli Viloria, L. Miller & J. Miller, 2008
SUBS Neopedaliodes michaeli innupta Pyrcz, 2008
SUBS Neopedaliodes michaeli citrica Viloria, L. Miller & J. Miller, 2008

SPEC Neopedaliodes proculeja Thieme, 1905

SUBS Neopedaliodes proculeja malaga Pyrcz, 2008
SUBS Neopedaliodes proculeja proculeja Thieme, 1905

SPEC Neopedaliodes phoenicusa (Hewitson, 1868)
SPEC Neopedaliodes yeyo Pyrcz & Rodriguez, 2008
SPEC Neopedaliodes tamaensis (Viloria & Pyrcz, 2007)

GENU Pedaliodes Butler, 1867

SPEC Pedaliodes adamsi D'Abrera, 1988
SPEC Pedaliodes albicilia Pyrcz, 2004
SPEC Pedaliodes albutia Thieme, 1905
SPEC Pedaliodes alusana (Hewitson, 1869)
SPEC Pedaliodes amafania Thieme, 1905
SPEC Pedaliodes anchiphilonis Hayward, 1968
SPEC Pedaliodes antulla Thieme, 1905
SPEC Pedaliodes arnotti Adams, 1986
SPEC Pedaliodes arturi Pyrcz & Viloria, 1999
SPEC Pedaliodes asconia Thieme, 1905
SPEC Pedaliodes auraria Thieme, 1905
SPEC Pedaliodes aureola Pyrcz, 2004
SPEC Pedaliodes auristriga Thieme, 1905
SPEC Pedaliodes baccara Thieme, 1905

SUBS Pedaliodes baccara baccara Thieme, 1905
SUBS Pedaliodes baccara allopatra Viloria & Pyrcz, 2007

SPEC Pedaliodes bernardi Adams, 1986
SPEC Pedaliodes boettgeri Pyrcz, 2004
SPEC Pedaliodes caeca Pyrcz & Viloria, 1999
SPEC Pedaliodes cebolleta Adams & Bernard, 1977
SPEC Pedaliodes cesarense Adams & Bernard, 1979
SPEC Pedaliodes chrysotaenia (Hopffer, 1874)
SPEC Pedaliodes cledonia Thieme, 1905
SPEC Pedaliodes coca Staudinger, 1894
SPEC Pedaliodes costipunctata Weymer, 1912
SPEC Pedaliodes daulis Thieme, 1905
SPEC Pedaliodes demathani Pyrcz, 2004

SUBS Pedaliodes demathani demathani Pyrcz, 2004
SUBS Pedaliodes demathani pisca Lamas & Viloria, 2008
SUBS Pedaliodes demathani kuni Pyrcz & Boyer, 2008
SUBS Pedaliodes demathani chaneli Pyrcz & Boyer, 2008
SUBS Pedaliodes demathani benedeki Pyrcz, 2008
SUBS Pedaliodes demathani alfa Pyrcz & Boyer, 2008
SUBS Pedaliodes demathani ockendeni Lamas & Viloria, 2008
SUBS Pedaliodes demathani kalinowskii Pyrcz & Boyer, 2008
SUBS Pedaliodes demathani horsti Pyrcz & Viloria, 2008
SUBS Pedaliodes demathani nussi Pyrcz, 2008

SPEC Pedaliodes derisoris Pyrcz, Viloria & Lamas, 2008
SPEC Pedaliodes dracula Pyrcz & Viloria, 1999
SPEC Pedaliodes empusa (C. Felder & R. Felder, 1867)

SUBS Pedaliodes empusa empusa (C. Felder & R. Felder, 1867)
SUBS Pedaliodes empusa medusa Pyrcz & Viloria, 2007

SPEC Pedaliodes erschoffi Pyrcz, 2004
SPEC Pedaliodes exanima (Erschoff, 1875)
SPEC Pedaliodes fassli Weymer, 1912
SPEC Pedaliodes ferratilis Butler, 1873
SPEC Pedaliodes fuscata (C. Felder & R. Felder, 1867)
SPEC Pedaliodes guicana Adams, 1986
SPEC Pedaliodes gustavi Viloria, L. Miller & J. Miller, 2003
SPEC Pedaliodes hardyi Adams, 1986
SPEC Pedaliodes hebena Pyrcz & Viloria, 1999
SPEC Pedaliodes hewitsoni Staudinger, 1897

SUBS Pedaliodes hewitsoni hewitsoni Staudinger, 1897
SUBS Pedaliodes hewitsoni primera Weeks, 1901

SPEC Pedaliodes hopfferi Staudinger, 1887
SPEC Pedaliodes japhleta Butler, 1870
SPEC Pedaliodes jelskii Pyrcz, 2004
SPEC Pedaliodes leucocheilus Godman & Salvin, 1880
SPEC Pedaliodes luperca Thieme, 1905
SPEC Pedaliodes maruda Pyrcz, 2004
SPEC Pedaliodes melaleuca Weymer, 1890
SPEC Pedaliodes misericordiosa Pyrcz & Viloria, 2008
SPEC Pedaliodes molesta Pyrcz, 2004
SPEC Pedaliodes morenoi Dognin, 1887

SUBS Pedaliodes morenoi morenoi Dognin, 1887
SUBS Pedaliodes morenoi pilaloensis Pyrcz & Viloria, 1999

SPEC Pedaliodes negreti Pyrcz, 1999
SPEC Pedaliodes niveonota Butler, 1873
SPEC Pedaliodes obscura Grose-Smith & W. F. Kirby, 1894
SPEC Pedaliodes obstructa Pyrcz & Viloria, 1999
SPEC Pedaliodes occulta Pyrcz & Viloria, 1999
SPEC Pedaliodes ochrotaenia (C. Felder & R. Felder, 1867)
SPEC Pedaliodes ornata Grose-Smith & W. F. Kirby, 1895
SPEC Pedaliodes pacifica E. Krüger, 1924
SPEC Pedaliodes pactyes (Hewitson, 1874)
SPEC Pedaliodes paeonides (Hewitson, 1874)
SPEC Pedaliodes palaepolis (Hewitson, 1878)
SPEC Pedaliodes pallantis (Hewitson, 1862)
SPEC Pedaliodes palpita Adams, 1986

SUBS Pedaliodes palpita palpita Adams, 1986
SUBS Pedaliodes palpita bedonica Pyrcz, 1999

SPEC Pedaliodes pammenes (Hewitson, 1874)
SPEC Pedaliodes paneis (Hewitson, 1862)
SPEC Pedaliodes panthides (Hewitson, 1874)
SPEC Pedaliodes parma Thieme, 1905
SPEC Pedaliodes parranda Adams, 1986
SPEC Pedaliodes patizathes (Hewitson, 1874)
SPEC Pedaliodes pausia (Hewitson, 1862)

SUBS Pedaliodes pausia pausia (Hewitson, 1862)
SUBS Pedaliodes pausia uncus Thieme, 1905

SPEC Pedaliodes pedacia (Hewitson, 1868)
SPEC Pedaliodes pelinaea (Hewitson, 1878)
SPEC Pedaliodes pelinna (Hewitson, 1870)
SPEC Pedaliodes perisades (Hewitson, 1874)
SPEC Pedaliodes peruda (Hewitson, 1862)
SPEC Pedaliodes peruviana Butler, 1873
SPEC Pedaliodes petri Pyrcz & Viloria, 1999

SUBS Pedaliodes petri petri Pyrcz & Viloria, 1999
SUBS Pedaliodes petri maasseni Pyrcz, 2004

SPEC Pedaliodes peucestas (Hewitson, 1862)

SUBS Pedaliodes peucestas peucestas (Hewitson, 1862)
SUBS Pedaliodes peucestas restricta Pyrcz & Viloria, 1999

SPEC Pedaliodes phaea (Hewitson, 1862)
SPEC Pedaliodes phaedra (Hewitson, 1870)

SUBS Pedaliodes phaedra phaedra (Hewitson, 1870)
SUBS Pedaliodes phaedra niphoessa Thieme, 1905

SPEC Pedaliodes phaeina Staudinger, 1897
SPEC Pedaliodes phanoclea (Hewitson, 1877)
SPEC Pedaliodes phazania Grose-Smith, 1900

SUBS Pedaliodes phazania phazania Grose-Smith, 1900
SUBS Pedaliodes phazania takahashii Adams & Bernard, 1977

SPEC Pedaliodes pheres Thieme, 1905
SPEC Pedaliodes pheretias (Hewitson, 1872)
SPEC Pedaliodes philonis (Hewitson, 1872)
SPEC Pedaliodes phoenissa (Hewitson, 1862)
SPEC Pedaliodes phoenix Lamas, 1999
SPEC Pedaliodes phrasa Grose-Smith & W. F. Kirby, 1894
SPEC Pedaliodes phrasicla (Hewitson, 1874)

SUBS Pedaliodes phrasicla phrasicla (Hewitson, 1874)
SUBS Pedaliodes phrasicla galaxias Weymer, 1912
SUBS Pedaliodes phrasicla immaculata Pyrcz, 1999
SUBS Pedaliodes phrasicla rufa Pyrcz, 2004

SPEC Pedaliodes phthiotis (Hewitson, 1874)
SPEC Pedaliodes piletha (Hewitson, 1862)
SPEC Pedaliodes piscolabis Pyrcz, 1999
SPEC Pedaliodes plotina (Hewitson, 1862)

SUBS Pedaliodes plotina plotina (Hewitson, 1862)
SUBS Pedaliodes plotina lucipara Weymer, 1912
SUBS Pedaliodes plotina pharnaspes (Hewitson, 1874)
SUBS Pedaliodes plotina rapha Butler, 1870
SUBS Pedaliodes plotina ruricola Pyrcz & Viloria, 2007

SPEC Pedaliodes poema Pyrcz & Viloria, 1999
SPEC Pedaliodes poesia (Hewitson, 1862)
SPEC Pedaliodes polla Thieme, 1905
SPEC Pedaliodes pollonia Adams, 1986
SPEC Pedaliodes polusca (Hewitson, 1862)
SPEC Pedaliodes porcia (Hewitson, 1869)

SUBS Pedaliodes porcia porcia (Hewitson, 1869)
SUBS Pedaliodes porcia pallantias (Hewitson, 1874)

SPEC Pedaliodes porima Grose-Smith & W. F. Kirby, 1894
SPEC Pedaliodes porina (Hewitson, 1862)
SPEC Pedaliodes praemontagna Pyrcz & Viloria, 2007
SPEC Pedaliodes praxia (Hewitson, 1877)
SPEC Pedaliodes praxithea (Hewitson, 1870)
SPEC Pedaliodes proerna (Hewitson, 1862)

SUBS Pedaliodes proerna proerna (Hewitson, 1862)
SUBS Pedaliodes proerna fumaria Thieme, 1905

SPEC Pedaliodes prytanis (Hewitson, 1862)
SPEC Pedaliodes puracana E. Krüger, 1924
SPEC Pedaliodes pylas (Hewitson, 1862)
SPEC Pedaliodes ralphi Adams, 1986
SPEC Pedaliodes reyi Pyrcz & Viloria, 2007
SPEC Pedaliodes rudnyi Pyrcz, 1999
SPEC Pedaliodes rumba Pyrcz & Viloria, 1999
SPEC Pedaliodes simmias Thieme, 1905
SPEC Pedaliodes simpla Thieme, 1905
SPEC Pedaliodes simplissima Pyrcz, 2004
SPEC Pedaliodes socorrae Adams, 1986
SPEC Pedaliodes sophismata Pyrcz, 2004
SPEC Pedaliodes spina Weymer, 1912
SPEC Pedaliodes stuebeli Pyrcz, 2004
SPEC Pedaliodes suspiro Adams & Bernard, 1979
SPEC Pedaliodes syleus Thieme, 1905
SPEC Pedaliodes symmachus Godman & Salvin, 1880
SPEC Pedaliodes sztolcmani Pyrcz, 2004

SUBS Pedaliodes sztolcmani sztolcmani Pyrcz, 2004
SUBS Pedaliodes sztolcmani gilvaecosta Pyrcz, 2004

SPEC Pedaliodes tabaconas Pyrcz & Viloria, 1999
SPEC Pedaliodes thiemei Staudinger, 1897
SPEC Pedaliodes transmontana Pyrcz & Viloria, 1999
SPEC Pedaliodes tucca Thieme, 1905
SPEC Pedaliodes tyro Thieme, 1905
SPEC Pedaliodes tyrrheoides Adams & Bernard, 1979
SPEC Pedaliodes tyrrheus Godman & Salvin, 1880

SUBS Pedaliodes tyrrheus tyrrheus Godman & Salvin, 1880
SUBS Pedaliodes tyrrheus tairona Adams & Bernard, 1977

SPEC Pedaliodes uaniuna Pyrcz & Viloria, 2004
SPEC Pedaliodes uniformis Weymer, 1912
SPEC Pedaliodes uniplaga Weymer, 1912
SPEC Pedaliodes valencia Pyrcz & Viloria, 2007
SPEC Pedaliodes vallenata Adams & Bernard, 1979
SPEC Pedaliodes wilhelmi Pyrcz, 1999
SPEC Pedaliodes woytkowskii Pyrcz, 2004
SPEC Pedaliodes xanthosphenisca Hayward, 1968
SPEC Pedaliodes zuleta Adams & Bernard, 1979
SPEC Pedaliodes haydoni Pyrcz & Viloria, 2013
SPEC Pedaliodes adrianae Pyrcz & Prieto, 2013
SPEC Pedaliodes rodriguezi Pyrcz & Andrade, 2013
SPEC Pedaliodes philinae Pyrcz & Prieto, 2013
SPEC Pedaliodes cocytia (C. Felder & R. Felder, 1867)
SPEC Pedaliodes nebris Thieme, 1905

SSYN Altopedaliodes similis Henao, Páez and Rodríguez-M., 2017

GENU Protopedaliodes Viloria & Pyrcz, 1994

SPEC Protopedaliodes kukenani Viloria & Pyrcz, 1994
SPEC Protopedaliodes ridouti Viloria & Pyrcz, 2000
SPEC Protopedaliodes profauna Viloria & Pyrcz, 2000
SPEC Protopedaliodes arekuna Pyrcz & Stachowicz, 2021

GENU Viloriodes Pyrcz & Espeland, 2022

SPEC Viloriodes manis (C. Felder & R. Felder, 1867)
SPEC Viloriodes ivica (Viloria & Pyrcz, 2010)
SPEC Viloriodes croizatorum (Viloria & Camacho, 1999)
SPEC Viloriodes antigua (Adams & Bernard, 1981)
SPEC Viloriodes circumducta (Thieme, 1905)
SPEC Viloriodes cremera (Godman & Salvin, 1878)
SPEC Viloriodes dejecta (H. Bates, 1865)
SPEC Viloriodes napaea (H. Bates, 1865)

SUBS Viloriodes napaea napaea (H. Bates, 1865)
SUBS Viloriodes napaea naksi (Pyrcz, 2012)

SPEC Viloriodes lithochalcis (Butler & Druce, 1872)
SPEC Viloriodes ereiba (C. Felder & R. Felder, 1867)
SPEC Viloriodes pomponia (Hewitson, 1869)
SPEC Viloriodes peregrina (Pyrcz & Viloria, 2012)
SPEC Viloriodes balnearia (Pyrcz & Viloria, 1999)
SPEC Viloriodes manneja (Thieme, 1905)
SPEC Viloriodes montagna (Adams & Bernard, 1981)
SPEC Viloriodes pisonia (Hewitson, 1862)
SPEC Viloriodes canela (Pyrcz & Viloria, 1999)
SPEC Viloriodes phrasis (Grose-Smith, 1900)
SPEC Viloriodes prosa (Staudinger, 1894)
SPEC Viloriodes phrasiclea (Grose-Smith, 1900)
SPEC Viloriodes pimienta (Adams, 1986)
SPEC Viloriodes zingara (Heredia & Viloria , 2004)
SPEC Viloriodes sonata (Pyrcz & Viloria, 1999)
SPEC Viloriodes roraimae (Strand, 1912)
SPEC Viloriodes chaconi (Viloria, 1998)
SPEC Viloriodes demarmelsi (Viloria, 1995)
SPEC Viloriodes terramaris (Viloria & Pyrcz, 2000)
SPEC Viloriodes triaria (Godman & Salvin, 1878)
SPEC Viloriodes yutajeana (Viloria & Pyrcz, 2000)
SPEC Viloriodes parakana (Viloria & Costa, 2019)

GENU Argyrophorus Blanchard, 1852

GSYN Stuardosatyrus Herrera & Etcheverry 1965
GSYN Etcheverrius Herrera 1966
GSYN Palmaris Herrera 1965
GSYN Pamperis Heimlich 1959
GSYN Punargentus Heimlich 1963

SPEC Argyrophorus argenteus Blanchard, 1852
SPEC Argyrophorus blanchardi Pyrcz & Wojtusiak, 2010

SUBS Argyrophorus blanchardi blanchardi Pyrcz & Wojtusiak, 2010
SUBS Argyrophorus blanchardi celendeni Pyrcz & Wojtusiak, 2010

SPEC Argyrophorus chiliensis (Guérin-Méneville, [1830])

SUBS Argyrophorus chiliensis chiliensis (Guérin-Méneville, [1830])
SUBS Argyrophorus chiliensis magallanicus Herrera, 1965
SUBS Argyrophorus chiliensis elwesi (Bryk, 1944)

SPEC Argyrophorus tandilensis (Köhler, 1935)
SPEC Argyrophorus antarcticus (Mabille, 1885)
SPEC Argyrophorus gustavi (Staudinger, 1898)

SUBS Argyrophorus gustavi gustavi (Staudinger, 1898)
SUBS Argyrophorus gustavi penai (Hayward, 1967)

SPEC Argyrophorus monticolens (A. Butler, 1881)
SPEC Argyrophorus poaoeneis (Heimlich, 1959)
SPEC Argyrophorus angusta Weymer, 1911

SUBS Argyrophorus angusta angusta Weymer, 1911
SUBS Argyrophorus angusta picota Fuchs, 1954

SPEC Argyrophorus lamna Thieme, 1904
SPEC Argyrophorus williamsianus Butler, 1868
SPEC Argyrophorus idealis Pyrcz & Cerdeña, 2019
SPEC Argyrophorus rubrostriata Pyrcz, Cerdeña & Florczyk, 2019

GENU Auca Hayward, 1953

SPEC Auca barrosi (C. Silva, 1917)
SPEC Auca coctei (Guérin-Méneville, [1838])
SPEC Auca pales (Philippi, 1859)

GENU Cosmosatyrus C. Felder & R. Felder, 1867

SPEC Cosmosatyrus leptoneuroides C. Felder & R. Felder, 1867

SUBS Cosmosatyrus leptoneuroides leptoneuroides C. Felder & R. Felder, 1867
SUBS Cosmosatyrus leptoneuroides plumbeola (Butler, 1868)

GENU Elina Blanchard, 1852

SPEC Elina montrolii (Feisthamel, 1839)
SPEC Elina vanessoides Blanchard, 1852

GENU Faunula C. Felder & R. Felder, 1867

SPEC Faunula leucoglene C. Felder & R. Felder, 1867
SPEC Faunula eleates (Weymer, 1890)
SPEC Faunula patagonica (Mabille, 1885)
SPEC Faunula dubii Pyrcz, 2012
SPEC Faunula euripides (Weymer, 1890)

SUBS Faunula euripides euripides (Weymer, 1890)
SUBS Faunula euripides salomonis (Köhler, 1939)

GENU Haywardella Herrera, 1966

SPEC Haywardella edmondsii (A. Butler, 1881)

GENU Homoeonympha C. Felder & R. Felder, 1867

GSYN Stygnus Felder & Felder 1867
GSYN Stygnolepis Strand 1942
GSYN Erebina Bryk 1944

SPEC Homoeonympha boisduvalii (Blanchard, 1852)

SUBS Homoeonympha boisduvalii boisduvalii (Blanchard, 1852)
SUBS Homoeonympha boisduvalii pusilla C. Felder & R. Felder, 1867

SPEC Homoeonympha vesagus (E. Doubleday, [1849])

GENU Nelia Hayward, 1953

SPEC Nelia calvertii (Elwes, 1903)
SPEC Nelia nemyroides (Blanchard, 1852)

GENU Neomaenas Wallengren, 1858

GSYN Stibomorpha Butler 1874

SPEC Neomaenas coenonymphina Butler, 1881
SPEC Neomaenas edmondsii (Butler, 1881)
SPEC Neomaenas fractifascia Butler, 1881
SPEC Neomaenas inornata (Elwes, 1903)
SPEC Neomaenas poliozona (C. Felder & R. Felder, 1867)
SPEC Neomaenas servilia Wallengren, 1858
SPEC Neomaenas wallengrenii Butler, 1881
SPEC Neomaenas ambiorix Wallengren, 1858
SPEC Neomaenas simplex (Butler, 1881)
SPEC Neomaenas humilis (C. Felder & R. Felder, 1867)
SPEC Neomaenas schajovskoii (Hayward, 1954)

GENU Pampasatyrus Hayward, 1953

GSYN Pseudocercyonis Miller & Emmel 1971

SPEC Pampasatyrus glaucope (C. Felder & R. Felder, 1867)

SUBS Pampasatyrus glaucope glaucope (C. Felder & R. Felder, 1867)
SUBS Pampasatyrus glaucope boenninghauseni (Foetterle, 1902)

SPEC Pampasatyrus gyrtone (Berg, 1877)
SPEC Pampasatyrus quies (Berg, 1877)
SPEC Pampasatyrus reticulata (Weymer, 1907)
SPEC Pampasatyrus yacantoensis (Köhler, 1939)
SPEC Pampasatyrus nilesi (Weeks, 1902)

SUBS Pampasatyrus nilesi nilesi (Weeks, 1902)
SUBS Pampasatyrus nilesi johanna (Weymer, 1911)

GENU Quilaphoetosus Herrera, 1966

SPEC Quilaphoetosus janirioides (Blanchard, 1852)
SPEC Quilaphoetosus monachus (Blanchard, 1852)

SUBS Quilaphoetosus monachus monachus (Blanchard, 1852)
SUBS Quilaphoetosus monachus valdiviae (C. Felder & R. Felder, 1867)

GENU Spinantenna Hayward, 1953

SPEC Spinantenna tristis (Guérin-Méneville, [1830])

GENU Tetraphlebia C. Felder & R. Felder, 1867

SPEC Tetraphlebia germainii C. Felder & R. Felder, 1867

SUBS Tetraphlebia germainii germainii C. Felder & R. Felder, 1867
SUBS Tetraphlebia germainii argentina Hayward, 1962

GENU Chillanella Herrera, 1966

SPEC Chillanella stelligera (A. Butler, 1881)

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