Nymphalidae Systematics Group

Checklist of Palaearctic Nymphalidae species


The list below is based on:

  • Bozano, G. C. (1999: Satyridae part I. Pages 1-58 in Guide to the Butterflies of the Palearctic Region (G. C. Bozano, ed.) Omnes Artes, Milano).
  • Bozano, G. C. (2002: Satyrinae part III. Pages 1-71 in Guide to the Butterflies of the Palearctic Region (G. C. Bozano, ed.) Omnes Artes, Milano).
  • Della Bruno, C., E. Gallo, M. Lucarelli, and V. Sbordoni. (2002: Satyrinae part II. Pages 1-59 in Guide to the Butterflies of the Palearctic Region (G. C. Bozano, ed.) Omnes Artes, Milano.
  • Tolman (1997: Butterflies of Britain and Europe. HarperCollins Publishers, London). Click here for a list of European species
  • Tuzov et al. (1997: Guide to the butterflies of Russia and adjacent territories. Volume 1. Pensoft, Sofia - Moscow).
  • Tuzov et al. (2000: Guide to the butterflies of Russia and adjacent territories. Volume 2. Pensoft, Sofia - Moscow).
  • Tuzov, V. K. (2003: Nymphalidae part I. Pages 1-64 in Guide to the Butterflies of the Palearctic Region (G. C. Bozano, ed.) Omnes Artes, Milano).
  • I do not have species marked in red for my molecular work on the systematics of Nymphalidae. I would very much appreciate samples of all these species! If you would like to help, please have a look at my advice on how to collect specimens for molecular work and send me an e-mail!

    The 662 Palaearctic species in Nymphalidae are:

    Subfamily Apaturinae

    Apatura metis ?, ?
    Apatura iris (Linnaeus, 1758)
    Apatura ilia (Denis & Schiffermüller, 1775)
    Athymodes nycteis (Ménétriés, 1859)
    Euapatura mirza ?, ?
    Mimathyma schrenckii (Ménétriés, 1859)
    Sephisa princeps (Fixsen, 1887)
    Thaleropis ionia (Eversmann, 1851)

    Subfamily Charaxinae

    Charaxes jasius (Linnaeus, 1767)

    Subfamily Danainae

    Danaus plexippus (Linnaeus, 1758)
    Danaus chrysippus (Linnaeus, 1758)
    Parantica sita (Kollar, 1844)

    Subfamily Heliconiinae

    Argynnis sagana Doubleday, 1847
    Argynnis childreni Gray, 1831
    Argynnis zenobia Leech, 1890
    Argynnis anadyomene Felder & Felder, 1862
    Argynnis ruslana Motschulsky, 1866
    Argynnis clara Blanchard, 1844
    Argynnis auresiana Fruhstorfer, 1908
    Argynnis jainadeva Moore, 1864
    Argynnis kamala Moore, 1857
    Argynnis coredippe Leech, 1893
    Argynnis xipe Grum-Grshimailo, 1891
    Argynnis nerippe Felder & Felder, 1862
    Argynnis elisa Godart, 1823
    Argynnis argyrospilata Kotzsch, 1938
    Argynnis vorax Butler, 1871
    Argynnis paphia (Linnaeus, 1758)
    Argynnis pandora (Denis & Schiffermüller, 1775)
    Argynnis laodice (Pallas, 1771)
    Argynnis aglaja (Linnaeus, 1758)
    Argynnis alexandra (Ménétriés, 1832)
    Argynnis adippe (Denis & Schiffermüller, 1775)
    Argynnis niobe (Linnaeus, 1758)
    Argynnis hyberbius (Linnaeus, 1763)
    Boloria pales ?, ?
    Boloria napaea ?, ?
    Boloria graeca ?, ?
    Boloria banghaasi Seitz, 1909
    Boloria purpurea Churkin, 1999
    Boloria frigidalis Warren, 1944
    Boloria aquilonaris (Stichel, 1908)
    Boloria caucasica (Lederer, 1852)
    Boloria sipora (Moore, 1875)
    Boloria altaica (Grum-Grshimailo, 1893)
    Boloria alaskensis (Holland, 1900)
    Boloria eunomia (Esper, 1799)
    Boloria angarensis (Erschoff, 1870)
    Boloria selene (Denis & Schiffermüller, 1775)
    Boloria oscarus (Eversmann, 1844)
    Boloria thore (Hübner, 1806)
    Boloria euphrosyne (Linnaeus, 1758)
    Boloria selenis (Eversmann, 1837)
    Boloria iphigenia (Graesser, 1888)
    Boloria perryi (Butler, 1882)
    Boloria hegemone (Staudinger, 1881)
    Boloria titania (Esper, 1793)
    Boloria dia (Linnaeus, 1767)
    Boloria butleri (Edwards, 1883)
    Boloria chariclea (Schneider, 1794)
    Boloria freija (Thunberg, 1791)
    Boloria polaris (Boisduval, 1828)
    Boloria frigga (Thunberg, 1791)
    Boloria improba (Butler, 1877)
    Boloria erda (Christoph, 1893)
    Boloria alberta (Edwards, 1890)
    Boloria tritonia (Boeber, 1812)
    Boloria distincta (Gibson, 1920)
    Boloria matveevi (Gorbunov & Korshunov, 1995)
    Brenthis mofidii Wyatt, 1969
    Brenthis hecate (Denis & Schiffermüller, 1775)
    Brenthis daphne (Bergsträsser, 1780)
    Brenthis ino (Rottemburg, 1775)
    Issoria lathonia (Linnaeus, 1758)
    Issoria eugenia (Eversmann, 1847)
    Issoria gemmata (Butler, 1881)
    Issoria altissima (Elwes, 1882)

    Subfamily Libytheinae

    Libythea celtis (Fuessly, 1782)

    Subfamily Limenitidinae

    Aldania raddei (Bremer, 1861)
    Aldania ilos (Fruhstorfer, 1909)
    Aldania themis (Leech, 1890)
    Aldania thisbe (Ménétriés, 1859)
    Aldania deliquata (Stichel, 1909)
    Limenitis reducta Staudinger, 1901
    Limenitis sidyi Lederer, 1853
    Limenitis moltrechti Kardakov, 1928
    Limenitis doerriesi Staudinger, 1892
    Limenitis homeyeri Tancré, 1881
    Limenitis helmanni Lederer, 1853
    Limenitis lepechini Erschoff, 1874
    Limenitis populi (Linnaeus, 1758)
    Limenitis camilla (Linnaeus, 1764)
    Limenitis amphyssa (Ménétriés, 1859)
    Neptis philyroides Staudinger, 1887
    Neptis speyeri Staudinger, 1887
    Neptis pryeri Butler, 1871
    Neptis alwina Bremer & Grey, 1852
    Neptis rivularis (Scopoli, 1763)
    Neptis sappho (Pallas, 1771)
    Neptis philyra (Ménétriés, 1859)
    Seokia pratti (Leech, 1890)

    Subfamily Nymphalinae

    Aglais io ?, ?
    Aglais urticae ?, ?
    Aglais caschmirensis ?, ?
    Aglais ladakhensis ?, ?
    Araschnia levana ?, ?
    Araschnia burejana ?, ?
    Euphydryas maturna ?, ?
    Euphydryas ichnea ?, ?
    Euphydryas iduna ?, ?
    Euphydryas cynthia ?, ?
    Euphydryas aurinia ?, ?
    Euphydryas asiatica ?, ?
    Euphydryas merope ?, ?
    Euphydryas orientalis ?, ?
    Euphydryas desfontainii ?, ?
    Hypolimnas misippus ?, ?
    Melitaea didyma ?, ?
    Melitaea latonigena ?, ?
    Melitaea interrupta ?, ?
    Melitaea persea ?, ?
    Melitaea saxatilis ?, ?
    Melitaea deserticola ?, ?
    Melitaea abyssinica ?, ?
    Melitaea casta ?, ?
    Melitaea didymina ?, ?
    Melitaea didymoides ?, ?
    Melitaea ala ?, ?
    Melitaea kotshubeji ?, ?
    Melitaea ninae ?, ?
    Melitaea chitralensis ?, ?
    Melitaea permuta ?, ?
    Melitaea enarea ?, ?
    Melitaea sutschana ?, ?
    Melitaea lutko ?, ?
    Melitaea mimetica ?, ?
    Melitaea acreina ?, ?
    Melitaea lunulata ?, ?
    Melitaea cassandra ?, ?
    Melitaea fergana ?, ?
    Melitaea infernalis ?, ?
    Melitaea ambrisia ?, ?
    Melitaea alraschid ?, ?
    Melitaea shandura ?, ?
    Melitaea athene ?, ?
    Melitaea trivia ?, ?
    Melitaea yuenty ?, ?
    Melitaea romanovi ?, ?
    Melitaea arduinna ?, ?
    Melitaea avinovi ?, ?
    Melitaea agar ?, ?
    Melitaea cinxia ?, ?
    Melitaea minerva ?, ?
    Melitaea balba ?, ?
    Melitaea sultanensis ?, ?
    Melitaea pallas ?, ?
    Melitaea turanica ?, ?
    Melitaea asteroida ?, ?
    Melitaea ludmilla ?, ?
    Melitaea solona ?, ?
    Melitaea arcesia ?, ?
    Melitaea leechi ?, ?
    Melitaea amoenula ?, ?
    Melitaea diamina ?, ?
    Melitaea protomedia ?, ?
    Melitaea phoebe ?, ?
    Melitaea scotosia ?, ?
    Melitaea telona ?, ?
    Melitaea punica ?, ?
    Melitaea sarvistana ?, ?
    Melitaea aetherie ?, ?
    Melitaea sibina ?, ?
    Melitaea pseudosibina ?, ?
    Melitaea collina ?, ?
    Melitaea turkmanica ?, ?
    Melitaea consulis ?, ?
    Melitaea gina ?, ?
    Melitaea athalia ?, ?
    Melitaea caucasogenita ?, ?
    Melitaea ambigua ?, ?
    Melitaea britomartis ?, ?
    Melitaea deione ?, ?
    Melitaea aurelia ?, ?
    Melitaea parthenoides ?, ?
    Melitaea varia ?, ?
    Melitaea asteria ?, ?
    Melitaea menetriesi ?, ?
    Melitaea rebeli ?, ?
    Melitaea westsibirica ?, ?
    Melitaea centralasiae ?, ?
    Melitaea plotina ?, ?
    Melitaea alatauica ?, ?
    Nymphalis antiopa ?, ?
    Nymphalis polychloros ?, ?
    Nymphalis xanthomelas ?, ?
    Nymphalis l-album ?, ?
    Polygonia c-album ?, ?
    Polygonia interposita ?, ?
    Polygonia egea ?, ?
    Polygonia canace ?, ?
    Vanessa atalanta ?, ?
    Vanessa vulcanica ?, ?
    Vanessa indica ?, ?
    Vanessa cardui ?, ?
    Vanessa virginiensis ?, ?

    Subfamily Satyrinae

    Aphantopus hyperantus ?, ?
    Arethusana arethusa (Denis & Schiffermüller, 1775)
    Argestina karta Riley, 1922
    Argestina waltoni (Elwes, 1906)
    Argestina inconstans (South, 1913)
    Argestina pomena (Evans, 1915)
    Berberia abdelkader ?, ?
    Berberia lambessanus ?, ?
    Boeberia parmenio (Boeber, 1809)
    Brintesia circe (Fabricius, 1775)
    Callerebia orixa Moore, 1872
    Callerebia annadina Watkins, 1927
    Callerebia bailey South, 1913
    Callerebia hybrida Butler, 1880
    Callerebia polyphemus (Oberthür, 1876)
    Callerebia annada (Moore, 1857)
    Callerebia scanda (Kollar, 1886)
    Callerebia tsirava (Evans, 1915)
    Callerebia nirmala (Moore, 1865)
    Chazara briseis ?, ?
    Chazara prieuri ?, ?
    Chazara heydenreichi ?, ?
    Chazara persephone ?, ?
    Chazara enervata ?, ?
    Chazara kaufmanni ?, ?
    Chazara sartha ?, ?
    Chazara staudingeri ?, ?
    Chazara eitschbergeri ?, ?
    Chazara rangontavica ?, ?
    Chazara bischoffi ?, ?
    Chonala laurae Bozano, 1999
    Chonala episcopalis (Oberthür, 1885)
    Chonala masoni (Elwes, 1882)
    Chonala miyatai (Koiwaya, 1996)
    Coenonympha symphita Lederer, 1870
    Coenonympha mangeri Bang-Haas, 1927
    Coenonympha nolckeni Erschoff, 1874)
    Coenonympha fettigii Oberthür, 1874
    Coenonympha vaucheri Blachier, 1905
    Coenonympha semenovi Alphéraky, 1887
    Coenonympha tullia (Müller, 1764)
    Coenonympha rhodopensis (Elwes, 1900)
    Coenonympha glycerion (Borkhausen, 1788)
    Coenonympha iphioides (Staudinger, 1870)
    Coenonympha mahometana (Alphéraky, 1888)
    Coenonympha sunbecca (Eversmann, 1843)
    Coenonympha amaryllis (Stoll, 1782)
    Coenonympha sinica (Alphéraky, 1888)
    Coenonympha hero (Linnaeus, 1761)
    Coenonympha arcania (Linnaeus, 1761)
    Coenonympha gardetta (de Prunner, 1798)
    Coenonympha orientalis (Rebel, 1910)
    Coenonympha arcanioides (Pierret, 1837)
    Coenonympha leander (Esper, 1784)
    Coenonympha mongolica (Alphéraky, 1881)
    Coenonympha dorus (Esper, 1782)
    Coenonympha austauti (Oberthür, 1881)
    Coenonympha corinna (Hübner, 1803)
    Coenonympha elbana (Staudinger, 1901)
    Coenonympha saadi (Kollar, 1850)
    Coenonympha pamphilus (Linnaeus, 1758)
    Coenonympha thyrsis (Freyer, 1845)
    Coenonympha oedippus (Fabricius, 1787)
    Erebia ligea ?, ?
    Erebia euryale ?, ?
    Erebia eriphyle ?, ?
    Erebia manto ?, ?
    Erebia claudina ?, ?
    Erebia flavofasciata ?, ?
    Erebia epiphron ?, ?
    Erebia orientalis ?, ?
    Erebia christi ?, ?
    Erebia pharte ?, ?
    Erebia melampus ?, ?
    Erebia sudetica ?, ?
    Erebia aethiops ?, ?
    Erebia triaria ?, ?
    Erebia embla ?, ?
    Erebia disa ?, ?
    Erebia medusa ?, ?
    Erebia polaris ?, ?
    Erebia alberganus ?, ?
    Erebia pluto ?, ?
    Erebia gorge ?, ?
    Erebia aethiopella ?, ?
    Erebia rhodopensis ?, ?
    Erebia mnestra ?, ?
    Erebia gorgone ?, ?
    Erebia epistygne ?, ?
    Erebia tyndarus ?, ?
    Erebia cassioides ?, ?
    Erebia hispania ?, ?
    Erebia nivalis ?, ?
    Erebia calcaria ?, ?
    Erebia ottomana ?, ?
    Erebia pronoe ?, ?
    Erebia melas ?, ?
    Erebia lefebvrei ?, ?
    Erebia scipio ?, ?
    Erebia stirius ?, ?
    Erebia styx ?, ?
    Erebia montana ?, ?
    Erebia zapateri ?, ?
    Erebia neoridas ?, ?
    Erebia oeme ?, ?
    Erebia meolans ?, ?
    Erebia palarica ?, ?
    Erebia pandrose ?, ?
    Erebia sthennyo ?, ?
    Erebia cyclopia ?, ?
    Erebia wanga ?, ?
    Erebia rossi ?, ?
    Erebia melancholica ?, ?
    Erebia neriene ?, ?
    Erebia niphonica ?, ?
    Erebia ajanensis ?, ?
    Erebia arctica ?, ?
    Erebia jeniseiensis ?, ?
    Erebia keferstreini ?, ?
    Erebia kindermanni ?, ?
    Erebia stubbendorfii ?, ?
    Erebia maurisius ?, ?
    Erebia pawlowskii ?, ?
    Erebia theano ?, ?
    Erebia haberhaueri ?, ?
    Erebia turanica ?, ?
    Erebia callias ?, ?
    Erebia iranica ?, ?
    Erebia graucasica ?, ?
    Erebia dabanensis ?, ?
    Erebia occulta ?, ?
    Erebia kozhantshikovi ?, ?
    Erebia fletcheri ?, ?
    Erebia anyuica ?, ?
    Erebia erynnyn ?, ?
    Erebia sachaensis ?, ?
    Erebia fasciata ?, ?
    Erebia discoidalis ?, ?
    Erebia progne ?, ?
    Erebia radians ?, ?
    Erebia sokolovi ?, ?
    Erebia eugenia ?, ?
    Erebia kalmuka ?, ?
    Erebia tianschanica ?, ?
    Erebia ocnus ?, ?
    Erebia sibo ?, ?
    Erebia meta ?, ?
    Erebia hewitsoni ?, ?
    Eugrumia herse (Grum-Grshimailo, 1891)
    Eugrumia discalis (South, 1913)
    Eugrumia koenigi (Goltz, 1938)
    Hemadara minorata Goltz, 1939
    Hemadara seitzi Goltz, 1939
    Hemadara albiligata Goltz, 1939
    Hemadara narasingha (Moore, 1857)
    Hemadara delavayi (Oberthür, 1891)
    Hemadara ruricola (Leech, 1890)
    Hemadara rurigena (Leech, 1890)
    Hipparchia hermione ?, ?
    Hipparchia ellena ?, ?
    Hipparchia neomiris ?, ?
    Hipparchia volgensis ?, ?
    Hipparchia cretica ?, ?
    Hipparchia christenseni ?, ?
    Hipparchia azorinus ?, ?
    Hipparchia caldeirense ?, ?
    Hipparchia miguelensis ?, ?
    Hipparchia mersina ?, ?
    Hipparchia leighebi ?, ?
    Hipparchia algirica ?, ?
    Hipparchia fatua ?, ?
    Hipparchia hansii ?, ?
    Hipparchia powelli ?, ?
    Hipparchia fidia ?, ?
    Hipparchia wyssii ?, ?
    Hipparchia fagi (Scopoli, 1763)
    Hipparchia syriaca (Staudinger, 1871)
    Hipparchia autonoe (Esper, 1783)
    Hipparchia semele (Linnaeus, 1758)
    Hipparchia pellucida (Stauder, 1924)
    Hipparchia statilinus (Hüfnagel, 1766)
    Hipparchia parisatis (Kollar, 1849)
    Hipparchia stulta (Staudinger, 1882)
    Hyponephele maroccana ?, ?
    Hyponephele lycaon ?, ?
    Hyponephele lupina ?, ?
    Hyponephele zuvandica ?, ?
    Hyponephele dzhungarica ?, ?
    Hyponephele przhewalskyi ?, ?
    Hyponephele pasimelas ?, ?
    Hyponephele lycaonoides ?, ?
    Hyponephele interposita ?, ?
    Hyponephele galtscha ?, ?
    Hyponephele davendra ?, ?
    Hyponephele brevistigma ?, ?
    Hyponephele korshunovi ?, ?
    Hyponephele issykkuli ?, ?
    Hyponephele comara ?, ?
    Hyponephele tenuistigma ?, ?
    Hyponephele argyrostigma ?, ?
    Hyponephele dysdora ?, ?
    Hyponephele tristis ?, ?
    Hyponephele prasolovi ?, ?
    Hyponephele murzini ?, ?
    Hyponephele jasavi ?, ?
    Hyponephele hilaris ?, ?
    Hyponephele fortambeka ?, ?
    Hyponephele rueckbeili ?, ?
    Hyponephele capella ?, ?
    Hyponephele amardaea ?, ?
    Hyponephele perplexa ?, ?
    Hyponephele glasunovi ?, ?
    Hyponephele naubidensis ?, ?
    Hyponephele huebneri ?, ?
    Hyponephele fusca ?, ?
    Hyponephele naricina ?, ?
    Hyponephele naricoides ?, ?
    Hyponephele haberhaueri ?, ?
    Hyponephele germana ?, ?
    Hyponephele maureri ?, ?
    Hyponephele shirazica ?, ?
    Hyponephele cadusia ?, ?
    Hyponephele cadusina ?, ?
    Hyponephele laeta ?, ?
    Hyponephele pamira ?, ?
    Hyponephele kirghisa ?, ?
    Hyponephele sheljuzhkoi ?, ?
    Hyponephele pseudokirgisa ?, ?
    Hyponephele rubriceps ?, ?
    Karanasa bolorica ?, ?
    Karanasa decolorata ?, ?
    Karanasa maureri ?, ?
    Karanasa josephi ?, ?
    Karanasa leechi ?, ?
    Karanasa wilkinsi ?, ?
    Karanasa talastauana ?, ?
    Karanasa praestans ?, ?
    Karanasa kirgisorum ?, ?
    Karanasa latifasciata ?, ?
    Karanasa puengeleri ?, ?
    Karanasa tancrei ?, ?
    Karanasa hoffmanni ?, ?
    Karanasa incerta ?, ?
    Karanasa straminea ?, ?
    Karanasa regeli ?, ?
    Karanasa kasakstana ?, ?
    Karanasa abramovi ?, ?
    Karanasa pamira ?, ?
    Karanasa alpherakyi ?, ?
    Kirinia roxelana (Cramer, 1777)
    Kirinia climene (Esper, 1783)
    Kirinia epimenides (Ménétriés, 1859)
    Kirinia epaminondas (Staudinger, 1887)
    Kirinia eversmanni (Eversmann, 1847)
    Lasiommata menava Moore, 1865
    Lasiommata maeroides Felder & Felder, 1867
    Lasiommata maerula Felder & Felder, 1867
    Lasiommata megera (Linnaeus, 1767)
    Lasiommata maera (Linnaeus, 1758)
    Lasiommata petropolitana (Fabricius, 1787)
    Lasiommata adrastoides (Bienert, 1870)
    Lasiommata felix (Warnecke, 1929)
    Lasiommata hindukushica (Wyatt & Omoto, 1966)
    Lasiommata majuscula (Leech, 1892)
    Lasiommata meadewaldoi (Rothschild, 1917)
    Lasiommata minuscula (Oberthür, 1923)
    Lasiommata paramegaera (Hübner, 1824)
    Lasiommata schakra (Kollar, 1844)
    Lethe bailey South, 1913
    Lethe hayashii Koiwaya, 1993
    Lethe hecate Leech, 1891
    Lethe laodamia Leech, 1891
    Lethe marginalis Motschulsky, 1860
    Lethe nigrifascia Leech, 1890
    Lethe proxima Leech, 1892
    Lethe yunnana D'Abrera, 1990
    Lethe argentata (Leech, 1891)
    Lethe callipteris (Butler, 1877)
    Lethe diana (Butler, 1866)
    Lethe dura (Marshall, 1882)
    Lethe gracilis (Oberthür, 1886)
    Lethe helle (Leech, 1891)
    Lethe jalaurida (de Niceville, 1880)
    Lethe manzorum (Poujade, 1884)
    Lethe ocellata (Poujade, 1885)
    Lethe oculatissima (Poujade, 1885)
    Lethe procne (Leech, 1891)
    Lethe shirozui (Sugiyama, 1997)
    Lethe sicelis (Hewitson, 1862)
    Lethe uemurai (Sugiyama, 1994)
    Lopinga eckweileri Görgner, 1990
    Lopinga achine (Scopoli, 1763)
    Lopinga catena (Leech, 1890)
    Lopinga deidamia (Eversmann, 1851)
    Lopinga dumetorum (Oberthür, 1886)
    Lopinga fulvescens (Alphéraky, 1889)
    Lopinga gerdae (Nordström, 1939)
    Lopinga nemorum (Oberthür, 1887)
    Loxerebia yukikoae Sugiyama, 1992
    Loxerebia martyr Watkins, 1927
    Loxerebia saxicola (Oberthür, 1876)
    Loxerebia albipuncta (Leech, 1890)
    Loxerebia pratorum (Oberthür, 1886)
    Loxerebia carola (Oberthür, 1893)
    Loxerebia loczy (Frivaldszky, 1866)
    Loxerebia bocki (Oberthür, 1893)
    Loxerebia sylvicola (Moore, 1857)
    Loxerebia megalops (Alphéraky, 1895)
    Loxerebia innupta (South, 1913)
    Loxerebia phyllis (Leech, 1891)
    Loxerebia yphtimoides (Oberthür, 1891)
    Lyela myops (Staudinger, 1881)
    Maniola jurtina ?, ?
    Maniola megala ?, ?
    Maniola chia ?, ?
    Maniola nurag ?, ?
    Maniola telmessia ?, ?
    Maniola halicarnassus ?, ?
    Melanargia evartiane Wagener, 1976
    Melanargia grumi Standfuss, 1892
    Melanargia parce Staudinger, 1882
    Melanargia leda Leech, 1891
    Melanargia asiatica Oberthür & Houlbert, 1922
    Melanargia galathea (Linnaeus, 1758)
    Melanargia lucasi (Rambur, 1858)
    Melanargia lachesis (Hübner, 1790)
    Melanargia teneates (Ménétriés, 1832)
    Melanargia titea (Klug, 1832)
    Melanargia larissa (Geyer, 1828)
    Melanargia hylata (Ménétriés, 1832)
    Melanargia syriaca (Oberthür, 1894)
    Melanargia occitanica (Esper, 1793)
    Melanargia ines (Hoffmannsegg, 1804)
    Melanargia arge (Sulzer, 1776)
    Melanargia russiae (Esper, 1793)
    Melanargia halimede (Ménétriés, 1858)
    Melanargia meridionalis (Felder & Felder, 1862)
    Melanargia montana (Leech, 1890)
    Melanargia lugens (Honrath, 1888)
    Melanargia epimede (Staudinger, 1887)
    Melanargia ganymedes (Heyne, 1895)
    Minois dryas ?, ?
    Neope christi Oberthür, 1886
    Neope dejeani Oberthür, 1894
    Neope niphonica Butler, 1881
    Neope oberthuri Leech, 1891
    Neope shirozui Koiwaya, 1989
    Neope simulans Leech, 1891
    Neope agrestis (Oberthür, 1876)
    Neope armandii (Oberthür, 1876)
    Neope bremeri (Felder, 1862)
    Neope goschkevitschii (Ménétriés, 1859)
    Neope muirheadii (Felder, 1862)
    Neope pulaha (Moore, 1857)
    Neorina patria Leech, 1891
    Ninguta schrencki (Ménétriés, 1859)
    Oeneis norna ?, ?
    Oeneis bore ?, ?
    Oeneis glacialis ?, ?
    Oeneis aktashi ?, ?
    Oeneis oeno ?, ?
    Oeneis polixenes ?, ?
    Oeneis melissa ?, ?
    Oeneis ammon ?, ?
    Oeneis hora ?, ?
    Oeneis taygete ?, ?
    Oeneis alpina ?, ?
    Oeneis pansa ?, ?
    Oeneis tarpeia ?, ?
    Oeneis lederi ?, ?
    Oeneis sculda ?, ?
    Oeneis nanna ?, ?
    Oeneis brunhilda ?, ?
    Oeneis diluta ?, ?
    Oeneis urda ?, ?
    Oeneis elwesi ?, ?
    Oeneis mulla ?, ?
    Oeneis magna ?, ?
    Oeneis jutta ?, ?
    Paralasa ali Churkin & Tuzov, 1998
    Paralasa manioides Wyatt & Omoto, 1966
    Paralasa howarthi Sakai, 1978
    Paralasa horaki Tuzov, 1997
    Paralasa nepalica Paulus, 1983
    Paralasa shakti Wyatt, 1961
    Paralasa danorum Clench & Shoumatoff, 1956
    Paralasa hades (Staudinger, 1882)
    Paralasa nero (Staudinger, 1894)
    Paralasa afghana (Goltz, 1937)
    Paralasa shallada (Lang, 1880)
    Paralasa pagmanni (Bang-Haas, 1927)
    Paralasa maracandica (Erschoff, 1874)
    Paralasa mani (de Niceville, 1880)
    Paralasa jordana (Staudinger, 1882)
    Paralasa icelos (Grum-Grshimailo, 1891)
    Paralasa ida (Grum-Grshimailo, 1890)
    Paralasa kalinda (Moore, 1865)
    Paralasa chitralica (Evans, 1923)
    Paralasa pamira (Shchetkin, 1989)
    Paralasa yasina (Tytler, 1926)
    Paralasa kusnezovi (Avinov, 1910)
    Paralasa kotzschae (Goltz, 1937)
    Paralasa semenovi (Avinov, 1910)
    Paralasa langara (Shchetkin, 1991)
    Paralasa phantasta (Goltz, 1938)
    Paralasa nitida (Riley, 1922)
    Pararge xiphioides Staudinger, 1871
    Pararge aegeria (Linnaeus, 1758)
    Pararge xiphia (Fabricius, 1775)
    Proterebia afra (Fabricius, 1787)
    Pseudochazara atlantis ?, ?
    Pseudochazara graeca ?, ?
    Pseudochazara hippolyte ?, ?
    Pseudochazara geyeri ?, ?
    Pseudochazara mamurra ?, ?
    Pseudochazara orestes ?, ?
    Pseudochazara mniszechii ?, ?
    Pseudochazara cingovskii ?, ?
    Pseudochazara anthelea ?, ?
    Pseudochazara tshujaca ?, ?
    Pseudochazara euxina ?, ?
    Pseudochazara turkestana ?, ?
    Pseudochazara panjshira ?, ?
    Pseudochazara pakistana ?, ?
    Pseudochazara pelopea ?, ?
    Pseudochazara schahrudensis ?, ?
    Pseudochazara alpina ?, ?
    Pseudochazara guriensis ?, ?
    Pseudochazara nukatli ?, ?
    Pseudochazara beroe ?, ?
    Pseudochazara schakuhensis ?, ?
    Pseudochazara daghestana ?, ?
    Pseudochazara kanishka ?, ?
    Pseudochazara thelephassa ?, ?
    Pyronia tithonus ?, ?
    Pyronia cecilia ?, ?
    Pyronia bathseba ?, ?
    Pyronia janiroides ?, ?
    Rhaphicera moorei Butler, 1867
    Rhaphicera dumicola (Oberthür, 1876)
    Satyrus actaea ?, ?
    Satyrus ferula ?, ?
    Satyrus dryas ?, ?
    Satyrus stheno ?, ?
    Satyrus virbius ?, ?
    Satyrus amasinus ?, ?
    Satyrus iranicus ?, ?
    Satyrus effendi ?, ?
    Satyrus daubi ?, ?
    Satyrus nana ?, ?
    Satyrus pimpla ?, ?
    Satyrus orphei ?, ?
    Sinonympha amoena Lee, 1974
    Tatinga thibetana (Oberthür, 1876)
    Triphysa dohrnii Zeller, 1850
    Triphysa phryne (Pallas, 1771)
    Ypthima amphithea (Ménétriés, 1859)
    Ypthima baldus (Fabricius, 1775)

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