Nymphalidae Systematics Group

The subfamily SATYRINAE

The mammoth subfamily Satyrinae comprises over 2000 species placed in 255 genera and 9 tribes, according to the latest version of the Classification of Nymphalidae. As currently delimited, the subfamily includes the tribes Morphini, Brassolini and Amathusiini, which have traditionally been placed in their own subfamily. However, the most recent study by Peña et al. (2006) shows clearly that the three tribes are within Satyrinae. The taxonomy of the subfamily is quite chaotic at the moment and needs to be revised, though the pioneering work of Miller in 1968 has helped much in creating some sort of order. The phylogenetic relationships of the subfamily are slowly being resolved through studies by the NSG.


Key publications for Satyrinae systematics:

  1. Brooks, C. J. 1950. A revision of the genus Tenaris Hübner (Lepidoptera: Amathusiidae). Trans. R. Ent. Soc. Lond. 101: 179-238.
  2. Jordan, K. 1924. On Hypocysta and some allied genera of Satyrinae (Lepidoptera: Rhopalocera) from New Guinea and the Solomon Islands. Novitates Zoologica 31: 279-297.
  3. Miller, L. D. 1968. The higher classification, phylogeny, and zoogeography of the Satyridae. Memmoirs of the American Entomological Society, Philadelphia.
  4. Parsons, M. J. 1999. The Butterflies of Papua New Guinea: Their Systematics and Biology. Academic Press, London.
  5. Peña, C., and G. Lamas. 2005. Revision of the butterfly genus Forsterinaria Gray, 1973 (Lepidoptera: Nymphalidae, Satyrinae). Revista peruana biologia 12:5-48.
  6. Peña, C., N. Wahlberg, E. Weingartner, U. Kodandaramaiah, S. Nylin, A. V. L. Freitas, and A. V. Z. Brower. 2006. Higher level phylogeny of Satyrinae butterflies (Lepidoptera: Nymphalidae) based on DNA sequence data. Molecular Phylogenetics and Evolution 40:29-49.
  7. Penz, C. M., and P. J. DeVries. 2002. Phylogenetic analysis of Morpho butterflies (Nymphalidae, Morphinae): implications for classification and natural history. American Museum Novitates 3374: 1-33.
  8. Weingartner, E., N. Wahlberg, and S. Nylin. 2006. Speciation in Pararge (Satyrinae: Nymphalidae) butterflies North Africa is the source of ancestral populations of all Pararge species. Systematic Entomology 31:621-632.

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