Argynnini

Niklas Wahlberg

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Updated 2020-11-20. Based on

Markku Savela's pages

Williams, M. C. 2018: Afrotropical Butterflies 17th Edition.

The Butterflies of America

Lamas, G. 2004: Atlas of Neotropical Lepidoptera. Checklist: Part 4A, Hesperioidea - Papilionoidea. Scientific Publishers, Gainsville, Florida.

Simonsen, T. J., Wahlberg, N., Brower, A. V. Z. & de Jong, R. 2006: Morphology, molecules and Fritillaries: approaching a stable phylogeny for Argynnini (Lepidoptera: Nymphalidae). Insect Systematics and Evolution 37:405-418

Wahlberg, N., Leneveu, J., Kodandaramaiah, U., Peña, C., Nylin, S., Freitas, A. V. L. & Brower, A. V. Z. 2009: Nymphalid butterflies diversify following near demise at the Cretaceous/Tertiary boundary. Proceedings of the Royal Society Series B Biological Sciences 276: 4295-4302

Simonsen, T. J., Wahlberg, N., Warren, A. D. & Sperling, F. A. H. 2010: The evolutionary history of Boloria (Lepidoptera: Nymphalidae): phylogeny, zoogeography and larval foodplant relationships. Systematics and Biodiversity 8: 513-529

de Moya, R. S., Savage, W. K., Tenney, C., Bao, X., Wahlberg, N. & Hill, R. I. 2017: Interrelationships and diversification of Argynnis Fabricius and Speyeria Scudder butterflies. Systematic Entomology 42: 635-649

TRIB Argynnini Swainson, 1833

GENU Pardopsis Trimen, 1887

SPEC Pardopsis punctatissima (Boisduval, 1833)

ORIG Acraea punctatissima Boisduval, 1833
SSYN Acraea stictica Boisduval, 1847

GENU Euptoieta Doubleday, 1848

SPEC Euptoieta bogotana Staudinger, 1885

SSYN Euptoieta colombia Strecker, 1899

SPEC Euptoieta claudia (Cramer, 1775)

SUBS Euptoieta claudia claudia (Cramer, 1775)

ORIG Papilio claudia Cramer, 1775
SSYN Papilio thais Doubleday, 1848

SUBS Euptoieta claudia daunius (Herbst, 1798)

ORIG Papilio daunius Herbst, 1798
SSYN Euptoieta claudia f. dodgei Gunder, 1927
SSYN Euptoieta claudia f. fumosa Field, 1936
SSYN Euptoieta claudia f. albaclaudia Field, 1936

SPEC Euptoieta hegesia (Cramer, 1779)

SUBS Euptoieta hegesia hegesia (Cramer, 1779)

ORIG Papilio hegesia Cramer, 1779
SSYN Euptoieta hegesia karibica Stichel, 1938
SSYN Euptoieta hegesia domingensis Cucurullo, 1959

SUBS Euptoieta hegesia meridiana Stichel, 1938

SSYN Euptoieta hegesia hoffmanni Comstock, 1944

SPEC Euptoieta hortensia (Blanchard, 1852)

ORIG Argynnis hortensia Blanchard, 1852
SSYN Argynnis valdiviana Philippi, 1859
SSYN Euptoieta ramirezi Giacomelli, 1926
SSYN Euptoieta claudia ab. infuscata Hayward, 1940

SPEC Euptoieta perdistincta Hall, 1930

SSYN Brenthis hana Clark, 1936

SPEC Euptoieta poasina Schaus, 1913
SPEC Euptoieta sunides (Hewitson, 1877)

ORIG Argynnis sunides Hewitson, 1877

SPEC Euptoieta thekla Hall, 1919

GENU Yramea Reuss, 1920

GSYN Chilargynnis Bryk, 1944

SPEC Yramea cytheris (Drury, 1773)

SUBS Yramea cyhteris cytheris (Drury, 1773)

ORIG Papilio cytheris Drury, 1773
SSYN Argynnis cytheris falklandica Watkins, 1924

SUBS Yramea cyhteris siga (Geyer, 1832)

ORIG Argynnis siga Geyer, 1832
SSYN Argynnis anna Blanchard, 1852
SSYN Argynnis chilensis Reed, 1877
SSYN Chilargynnis cytheris subtusviola Bryk, 1944

SPEC Yramea inca (Staudinger, 1894)

SUBS Yramea inca inca (Staudinger, 1894)

ORIG Argynnis inca Staudinger, 1894

SUBS Yramea inca cora (Lehmann, 1913)

ORIG Argynnis cora Lehmann, 1913

SPEC Yramea lathonioides (Blanchard, 1852)

ORIG Argynnis lathonioides Blanchard, 1852
SSYN Argynnis dexamene Boisduval, 1859
SSYN Argynnis dioides Boisduval, 1859
SSYN Argynnis montana Reed, 1877
SSYN Argynnis euterpe Reed, 1877
SSYN Argynnis darwini Staudinger, 1899

SPEC Yramea modesta (Blanchard, 1852)

ORIG Argynnis modesta Blanchard, 1852

SPEC Yramea sobrina (Weymer, 1890)

ORIG Argynnis sobrina Weymer, 1890
SSYN Boloria tessellata Rosenberg & Talbot, 1914
SSYN Yramea blancasi Hayward, 1960

SPEC Yramea lynx Lamas & Grados, 2004

GENU Argynnis Fabricius, 1807

GSYN Pandoriana Warren, 1942
GSYN Childrena Hemming, 1943
GSYN Eudryas Reuss, 1926 (preocc. Eudryas Boisduval, 1836; Eudryas Harris, 1841; Eudryas Fitzinger, 1843; Eudryas Gistl, 1848)
GSYN Argyreus Scopoli, 1777
GSYN Acidalia Hübner, 1819
GSYN Argyrea Billberg, 1820
GSYN Mimargyra Reuss, 1922
GSYN Damora Nordmann, 1851
GSYN Argyronome Hübner, 1819

SPEC Argynnis paphia (Linnaeus, 1758)

ORIG Papilio paphia Linnaeus, 1758

SUBS Argynnis paphia paphia (Linnaeus, 1758)
SUBS Argynnis paphia valesina Esper
SUBS Argynnis paphia dives (Oberthür, 1908)
SUBS Argynnis paphia formosicola Matsumura
SUBS Argynnis paphia neopaphia Fruhstorfer
SUBS Argynnis paphia megalegoria Fruhstorfer, 1907

SPEC Argynnis sagana Doubleday, 1847

SUBS Damora sagana sagana (Doubleday, 1847)
SUBS Damora sagana liane (Fruhstorfer, 1907)
SUBS Damora sagana relicta Korshunov, 1984
SUBS Damora sagana paulina (Nordman, 1851)

SPEC Argynnis anadyomene Felder & Felder, 1862
SPEC Argynnis pandora (Denis & Schiffermüller, 1775)

ORIG Papilio pandora Denis & Schiffermüller, 1775
SSYN Papilio maja Cramer, 1775
SSYN Pandoriana pandora (Denis & Schiffermüller, 1775)

SUBS Argynnis pandora pandora (Denis & Schiffermüller, 1775)
SUBS Argynnis pandora seitzi Fruhstorfer, 1908

SPEC Argynnis childreni Gray, 1831

SUBS Argynnis childreni childreni Gray, 1831
SUBS Argynnis childreni sakontala Kollar
SUBS Argynnis childreni caesarea Fruhstorfer

SPEC Argynnis zenobia (Leech, 1890)

SUBS Argynnis zenobia zenobia (Leech, 1890)
SUBS Argynnis zenobia penelope (Staudinger)

SPEC Argynnis hyperbius (Linnaeus, 1763)

ORIG Papilio hyperbius Linnaeus, 1763
SSYN Papilio niphe Linnaeus, 1767 (unnec. repl. for Papilio hyperbius Linnaeus, 1763)

SUBS Argynnis hyperbius hyperbius (Linnaeus, 1763)
SUBS Argynnis hyperbius taprobana Moore
SUBS Argynnis hyperbius castetsi Oberthür
SUBS Argynnis hyperbius sumatrensis (Fruhstorfer, 1905)
SUBS Argynnis hyperbius javanica Oberthür
SUBS Argynnis hyperbius sagada Fruhstorfer
SUBS Argynnis hyperbius inconstans (Butler, 1873)
SUBS Argynnis hyperbius neumanni Rothschild & Jordan

SPEC Argynnis laodice (Pallas, 1771)

ORIG Papilio laodice Pallas, 1771

SUBS Argynnis laodice laodice (Pallas, 1771)
SUBS Argynnis laodice rudra (Moore, 1857)
SUBS Argynnis laodice samana Fruhstorfer, 1902
SUBS Argynnis laodice japonica (Ménétriés)
SUBS Argynnis laodice huochengice Huang & Murayama

SPEC Argynnis kuniga Chou & Tong, 1994
SPEC Argynnis ruslana Motschulsky, 1866

GENU Speyeria Scudder, 1872

GSYN Semnopsyche Scudder, 1875
GSYN Mesoacidalia Reuss, 1926
GSYN Proacidalia Reuss, 1926

SPEC Speyeria aglaja (Linnaeus, 1758)

ORIG Papilio aglaja Linnaeus, 1758
SSYN Papilio pasilhoe Linnaeus, 1767
SSYN Papilio charlotta Haworth, 1802
SSYN Mesoacidalia aglaja (Linnaeus, 1758)

SUBS Speyeria aglaja aglaja (Linnaeus, 1758)
SUBS Speyeria aglaja lyauteyi (Oberthür, 1920)
SUBS Speyeria aglaja excelsior (Rothschild, 1933)
SUBS Speyeria aglaja vitatha (Moore)
SUBS Speyeria aglaja clavimacula (Matsumura)
SUBS Speyeria aglaja bessa (Fruhstorfer, 1907)

SPEC Speyeria clara (Blanchard, 1844)

ORIG Argynnis clara Blanchard, 1844

SPEC Speyeria alexandra Ménétriés, 1855
SPEC Speyeria claudia Fawcett, 1904
SPEC Speyeria diana (Cramer, 1777)

ORIG Papilio diana Cramer, 1777

SPEC Speyeria cybele (Fabricius, 1775)

ORIG Papilio cybele (Fabricius, 1775)

SUBS Speyeria cybele cybele (Fabricius, 1775)
SUBS Speyeria cybele novascotiae (McDunnough, 1935)
SUBS Speyeria cybele krautwurmi (W. Holland, 1931)
SUBS Speyeria cybele pseudocarpenteri (F. Chermock & R. Chermock, 1940)
SUBS Speyeria cybele carpenterii (W. H. Edwards, 1876)
SUBS Speyeria cybele charlottii (W. Barnes, 1897)
SUBS Speyeria cybele letona dos Passos & Grey, 1945
SUBS Speyeria cybele leto (Behr, 1862)
SUBS Speyeria cybele pugetensis F. Chermock & Frechin, 1947
SUBS Speyeria cybele eileenae J. Emmel, T. Emmel & Mattoon, 1998

SPEC Speyeria aphrodite (Fabricius, 1787) Aphrodite Fritillary

ORIG Papilio aphrodite (Fabricius, 1787)

SUBS Speyeria aphrodite cullasaja Gatrelle, 1998
SUBS Speyeria aphrodite aphrodite (Fabricius, 1787)
SUBS Speyeria aphrodite alcestis (W. H. Edwards, [1877])
SUBS Speyeria aphrodite ethne (Hemming, 1933)
SUBS Speyeria aphrodite byblis (W. Barnes & Benjamin, 1926)
SUBS Speyeria aphrodite columbia (H. Edwards, 1877)
SUBS Speyeria aphrodite whitehousei (Gunder, 1932)
SUBS Speyeria aphrodite manitoba (F. Chermock & R. Chermock, 1940)
SUBS Speyeria aphrodite winni (Gunder, 1932)

SPEC Speyeria idalia (Drury, 1773)

SUBS Speyeria idalia idalia (Drury, 1773)
SUBS Speyeria idalia occidentalis B. Williams, 2002

SPEC Speyeria nokomis (W. H. Edwards, 1862)

SUBS Speyeria nokomis coerulescens (W. Holland, 1900)
SUBS Speyeria nokomis nitocris (W. H. Edwards, 1874)
SUBS Speyeria nokomis nokomis (W. H. Edwards, 1862)
SUBS Speyeria nokomis tularosa (R. Holland, 2010)
SUBS Speyeria nokomis apacheana (Skinner, 1918)
SUBS Speyeria nokomis carsonensis Austin, 1998
SUBS Speyeria nokomis wenona dos Passos & Grey, 1945
SUBS Speyeria nokomis melaena Mooser & García, 1980

SPEC Speyeria edwardsii (Reakirt, 1866)
SPEC Speyeria coronis (Behr, 1864)

SUBS Speyeria coronis semiramis (W. H. Edwards, 1886)
SUBS Speyeria coronis hennei (Gunder, 1934)
SUBS Speyeria coronis coronis (Behr, 1864)
SUBS Speyeria coronis simaetha dos Passos & Grey, 1945
SUBS Speyeria coronis snyderi (Skinner, 1897)
SUBS Speyeria coronis halcyone (W. H. Edwards, [1869])

SPEC Speyeria carolae (dos Passos & Grey, 1942)
SPEC Speyeria zerene (Boisduval, 1852)

SUBS Speyeria zerene sitka Hammond, Harry & McCorkle, 2001
SUBS Speyeria zerene bremnerii (W. H. Edwards, 1872)
SUBS Speyeria zerene hippolyta (W. H. Edwards, 1879)
SUBS Speyeria zerene gloriosa Moeck, 1957
SUBS Speyeria zerene behrensii (W. H. Edwards, 1869)
SUBS Speyeria zerene sonomensis J. Emmel, T. Emmel & Mattoon, 1998
SUBS Speyeria zerene puntareyes J. Emmel & T. Emmel, 1998
SUBS Speyeria zerene myrtleae dos Passos & Grey, 1945
SUBS Speyeria zerene monticola (Behr, 1863)
SUBS Speyeria zerene malcolmi (J. A. Comstock, 1920)
SUBS Speyeria zerene zerene (Boisduval, 1852)
SUBS Speyeria zerene gunderi (J. A. Comstock, 1925)
SUBS Speyeria zerene picta (McDunnough, 1924)
SUBS Speyeria zerene platina (Skinner, 1897)
SUBS Speyeria zerene sinope dos Passos & Grey, 1945

SPEC Speyeria callippe (Boisduval, 1852)

SUBS Speyeria callippe adiasteoides J. Emmel, T. Emmel & Mattoon, 1998
SUBS Speyeria callippe liliana (H. Edwards, 1877)
SUBS Speyeria callippe callippe (Boisduval, 1852)
SUBS Speyeria callippe comstocki (Gunder, 1925)
SUBS Speyeria callippe macaria (W. H. Edwards, 1877)
SUBS Speyeria callippe laurina (W. G. Wright, 1905)
SUBS Speyeria callippe rupestris (Behr, 1863)
SUBS Speyeria callippe inornata (W. H. Edwards, 1872)
SUBS Speyeria callippe juba (Boisduval, 1869)
SUBS Speyeria callippe hagemani J. Emmel, T. Emmel & Mattoon, 1998
SUBS Speyeria callippe shasta J. Emmel, T. Emmel & Mattoon, 1998
SUBS Speyeria callippe elaine dos Passos & Grey, 1945
SUBS Speyeria callippe semivirida (McDunnough, 1924)
SUBS Speyeria callippe chilcotinensis C. Guppy & J. Shepard, 2001
SUBS Speyeria callippe nevadensis (W. H. Edwards, 1870)
SUBS Speyeria callippe harmonia dos Passos & Grey, 1945
SUBS Speyeria callippe calgariana (McDunnough, 1924)
SUBS Speyeria callippe gallatini (McDunnough, 1929)
SUBS Speyeria callippe meadii (W. H. Edwards, 1872)

SPEC Speyeria egleis (Behr, 1862)

SUBS Speyeria egleis tehachapina (J. A. Comstock, 1920)
SUBS Speyeria egleis egleis (Behr, 1862)
SUBS Speyeria egleis oweni (W. H. Edwards, 1892)
SUBS Speyeria egleis mattooni J. Emmel & T. Emmel, 1998
SUBS Speyeria egleis yolaboli J. Emmel & T. Emmel, 1998
SUBS Speyeria egleis moecki Hammond & Dornfeld, 1983
SUBS Speyeria egleis toiyabe Howe, 1975
SUBS Speyeria egleis reidi Austin, 1998
SUBS Speyeria egleis linda (dos Passos & Grey, 1942)
SUBS Speyeria egleis utahensis (Skinner, 1919)
SUBS Speyeria egleis secreta dos Passos & Grey, 1945
SUBS Speyeria egleis macdunnoughi (Gunder, 1932)
SUBS Speyeria egleis albrighti (Gunder, 1932)

SPEC Speyeria adiaste (W. H. Edwards, 1864)

SUBS Speyeria adiaste adiaste (W. H. Edwards, 1864)
SUBS Speyeria adiaste clemencei (J. A. Comstock, 1925)
SUBS Speyeria adiaste atossa (W. H. Edwards, 1890)

SPEC Speyeria atlantis (W. H. Edwards, 1862)

SUBS Speyeria atlantis atlantis (W. H. Edwards, 1862)
SUBS Speyeria atlantis canadensis (dos Passos, 1935)
SUBS Speyeria atlantis hollandi (F. Chermock & R. Chermock, 1940)
SUBS Speyeria atlantis pahasapa Spomer, Scott & Kondla, 1998
SUBS Speyeria atlantis sorocko Scott, Kondla & Spomer, 1998

SPEC Speyeria hesperis (W. H. Edwards, 1864)

SUBS Speyeria hesperis irene (Boisduval, 1869)
SUBS Speyeria hesperis hanseni J. Emmel, T. Emmel & Mattoon, 1998
SUBS Speyeria hesperis cottlei (J. A. Comstock, 1925)
SUBS Speyeria hesperis dodgei (Gunder, 1931)
SUBS Speyeria hesperis elko Austin, 1984
SUBS Speyeria hesperis greyi Moeck, 1950
SUBS Speyeria hesperis chitone (W. H. Edwards, 1879)
SUBS Speyeria hesperis schellbachi Garth, 1949
SUBS Speyeria hesperis tetonia dos Passos & Grey, 1945
SUBS Speyeria hesperis viola dos Passos & Grey, 1945
SUBS Speyeria hesperis hutchinsi dos Passos & Grey, 1947
SUBS Speyeria hesperis brico Kondla, Scott & Spomer, 1998
SUBS Speyeria hesperis beani (W. Barnes & Benjamin, 1926)
SUBS Speyeria hesperis dennisi dos Passos & Grey, 1945
SUBS Speyeria hesperis lurana dos Passos & Grey, 1945
SUBS Speyeria hesperis hesperis (W. H. Edwards, 1864)
SUBS Speyeria hesperis ratonensis Scott, 1981
SUBS Speyeria hesperis electa (W. H. Edwards, 1878)
SUBS Speyeria hesperis dorothea Moeck, 1947
SUBS Speyeria hesperis capitanensis R. Holland, 1988
SUBS Speyeria hesperis nausicaa (W. H. Edwards, 1874)

SPEC Speyeria hydaspe (Boisduval, 1869)

SUBS Speyeria hydaspe viridicornis (J. A. Comstock, 1925)
SUBS Speyeria hydaspe hydaspe (Boisduval, 1869)
SUBS Speyeria hydaspe tehama J. Emmel, T. Emmel & Mattoon, 1998
SUBS Speyeria hydaspe davisi J. Emmel, T. Emmel & Mattoon, 1998
SUBS Speyeria hydaspe rhodope (W. H. Edwards, 1874)
SUBS Speyeria hydaspe minor dos Passos & Grey, 1947

SPEC Speyeria mormonia (Boisduval, 1869)

SUBS Speyeria mormonia bischoffii (W. H. Edwards, 1870)
SUBS Speyeria mormonia opis (W. H. Edwards, 1874)
SUBS Speyeria mormonia washingtonia (W. Barnes & McDunnough, 1913)
SUBS Speyeria mormonia erinna (W. H. Edwards, 1883)
SUBS Speyeria mormonia kimimela (Marrone, Spomer & Scott, 2008)
SUBS Speyeria mormonia eurynome (W. H. Edwards, 1872)
SUBS Speyeria mormonia mormonia (Boisduval, 1869)
SUBS Speyeria mormonia obsidiana J. Emmel, T. Emmel & Mattoon, 1998
SUBS Speyeria mormonia artonis (W. H. Edwards, 1881)
SUBS Speyeria mormonia luski (W. Barnes & McDunnough, 1913)

GENU Fabriciana Reuss, 1920

GSYN Profabriciana Reuss, 1926
GSYN Protodryas Reuss, 1928

SPEC Fabriciana niobe (Linnaeus, 1758)

ORIG Papilio niobe Linnaeus, 1758

SUBS Fabriciana niobe niobe (Linnaeus, 1758)
SUBS Fabriciana niobe changaica Reuss, 1922
SUBS Fabriciana niobe ornata (Staudinger)
SUBS Fabriciana niobe orientalis (Alphéraky, 1881)

SPEC Fabriciana coredippe Leech, 1892
SPEC Fabriciana auresiana (Fruhstorfer, 1908)

ORIG Argynnis adippe auresiana Fruhstorfer, 1908

SUBS Fabriciana auresiana auresiana (Fruhstorfer, 1908)
SUBS Fabriciana auresiana hassani Weiss, 1979
SUBS Fabriciana auresiana astrifera Higgins, 1965

SPEC Fabriciana adippe (Denis & Schiffermüller, 1775)

ORIG Papilio adippe Denis & Schiffermüller, 1775
SSYN Papilio cydippe Linnaeus, 1761
SSYN Argynnis adippe (Denis & Schiffermüller, 1775)

SUBS Fabriciana adippe adippe (Denis & Schiffermüller, 1775)
SUBS Fabriciana adippe ornatissima (Leech, 1892)
SUBS Fabriciana adippe vorax (Butler, 1871)
SUBS Fabriciana adippe tianshanica (Staudinger)
SUBS Fabriciana adippe zarevna Fruhstorfer, 1912
SUBS Fabriciana adippe xanthodippe (Fixsen)
SUBS Fabriciana adippe taurica (Staudinger)
SUBS Fabriciana adippe taliana Reuss, 1922
SUBS Fabriciana adippe pallescens (Butler, 1873)
SUBS Fabriciana adippe chayuensis Huang, 2001

SPEC Fabriciana xipe (Groum-Grshimailo, 1891)

SUBS Fabriciana xipe xipe (Groum-Grshimailo, 1891)
SUBS Fabriciana xipe gyala (Tytler, 1940)

SPEC Fabriciana elisa (Godart, 1823)

ORIG Argynnis elisa Godart, 1823

SPEC Fabriciana argyrospilata Kotsch, 1938
SPEC Fabriciana nerippe (Felder, 1862)

SUBS Fabriciana nerippe nerippe (Felder, 1862)
SUBS Fabriciana nerippe nerippina (Fruhstorfer)

SPEC Fabriciana kamala (Moore, 1857)

ORIG Argynnis kamala Moore, 1857

SPEC Fabriciana jainadeva (Moore, 1864)

ORIG Argynnis jainadeva Moore, 1864
SSYN Profabriciana jainadeva (Moore, 1864)

SPEC Fabriciana funebris Oberthür

GENU Brenthis Hübner, 1819

SPEC Brenthis ino (Rottemburg, 1775)

ORIG Papilio ino (Rottemburg, 1775)

SUBS Brenthis ino ino (Rottemburg, 1775)
SUBS Brenthis ino paidicus Fruhstorfer, 1907
SUBS Brenthis ino acrita Fruhstorfer
SUBS Brenthis ino amurensis (Staudinger)
SUBS Brenthis ino discolus (Fruhstorfer)

SPEC Brenthis hecate (Denis & Schiffermüller, 1775)

ORIG Papilio hecate Denis & Schiffermüller, 1775

SUBS Brenthis hecate hecate (Denis & Schiffermüller, 1775)
SUBS Brenthis hecate alaica Staudinger
SUBS Brenthis hecate warren Kudrna, 1974

SPEC Brenthis mofidii Wyatt, 1969
SPEC Brenthis daphne (Denis & Schiffermüller, 1775)

ORIG Papilio daphne Denis & Schiffermüller, 1775

SUBS Brenthis daphne daphne (Denis & Schiffermüller, 1775)
SUBS Brenthis daphne ochroleuca (Fruhstorfer)

GENU Issoria Hübner, 1819

GSYN Rathora Moore, 1900
GSYN Prokuekenthaliella Reuss, 1927
GSYN Kuekenthaliella Reuss, 1921

SPEC Issoria lathonia (Linnaeus, 1758)

ORIG Papilio lathonia Linnaeus, 1758

SUBS Issoria lathonia lathonia (Linnaeus, 1758)
SUBS Issoria lathonia issaea (Gray, 1857)

SPEC Issoria hanningtoni (Elwes, 1889)

SUBS Issoria hanningtoni hanningtoni (Elwes, 1889)
SUBS Issoria hanningtoni jeanneli Bernardi, 1968
SUBS Issoria hanningtoni imatonga (Riley, 1932)

SPEC Issoria smaragdifera (Butler, 1895)

ORIG Argynnis smaragdifera Butler, 1895

SUBS Issoria smaragdifera smaragdifera (Butler, 1895)
SUBS Issoria smaragdifera reducta Carcasson, 1961

SPEC Issoria baumanni (Rebel & Rogenhofer, 1894)

SUBS Issoria baumanni baumanni (Rebel & Rogenhofer, 1894)
SUBS Issoria baumanni excelsior (Butler, 1896)

SSYN Argynnis uganda Röber, 1937

SUBS Issoria baumanni katangae (Neave, 1910)
SUBS Issoria baumanni orientalis Kielland, 1990

SPEC Issoria gemmata (Butler, 1881)

ORIG Argynnis gemmata Butler, 1881

SUBS Issoria gemmata gemmata (Butler, 1881)
SUBS Issoria gemmata wui Huang, 1998

SPEC Issoria baileyi Huang, 1998
SPEC Issoria eugenia (Eversmann, 1847)

SUBS Issoria eugenia eugenia (Eversmann, 1847)
SUBS Issoria eugenia montana (Bang-Haas, 1906)
SUBS Issoria eugenia rhea (Groum-Grshimailo, 1891)
SUBS Issoria eugenia genia (Fruhstorfer, 1903)
SUBS Issoria eugenia pulchella Huang, 2001
SUBS Issoria eugenia tibetana Huang, 1998

SPEC Issoria altissima (Elwes, 1882)
SPEC Issoria mackinnonii (de Nicéville, 1891)

GENU Boloria Moore, 1900

SPEC Boloria pales (Denis & Schiffermüller, 1775)

ORIG Papilio pales Denis & Schiffermüller, 1775

SUBS Boloria pales pales (Denis & Schiffermüller, 1775)
SUBS Boloria pales palina (Fruhstorfer, 1903)
SUBS Boloria pales eupales (Fruhstorfer, 1903)
SUBS Boloria pales shambhala Kocman, 1999
SUBS Boloria pales nirvana Kocman, 1999
SUBS Boloria pales sifanica (Grum-Grshimailo, 1891)

SPEC Boloria generator Staudinger, 1886
SPEC Boloria napaea (Hoffmannsegg, 1804)

SUBS Boloria napaea napaea (Hoffmannsegg, 1804)
SUBS Boloria napaea altaica (Groum-Grshimailo, 1893)
SUBS Boloria napaea alaskensis (Holland, 1900)
SUBS Boloria napaea nearctica Verity, 1932
SUBS Boloria napaea halli Klots, 1940

SPEC Boloria frigidalis Warren, 1944
SPEC Boloria aquilonaris (Stichel, 1980)

SUBS Boloria aquilonaris aquilonaris (Stichel, 1980)
SUBS Boloria aquilonaris banghaasi (Seitz, 1909)

SPEC Boloria graeca (Staudinger, 1870)
SPEC Boloria eunomia (Esper, 1799)

ORIG Papilio eunomia Esper, 1799
SSYN Papilio aphirape Hübner, 1799
SSYN Proclossiana eunomia (Esper, 1799)
SSYN Clossiana eunomia (Esper, 1799)

SUBS Boloria eunomia eunomia (Esper, 1799)
SUBS Boloria eunomia ossiana (Herbst, 1800)
SUBS Boloria eunomia triclaris (Hübner, 1821)
SUBS Boloria eunomia caelestis (Hemming, 1933)
SUBS Boloria eunomia dawsoni (Barnes & McDunnough, 1916)
SUBS Boloria eunomia nichollae (Barnes & Benjamin, 1926)
SUBS Boloria eunomia laddi (Klots, 1940)
SUBS Boloria eunomia ursadentis (Ferris & Groothuis, 1971)
SUBS Boloria eunomia denali (Klots, 1940)
SUBS Boloria eunomia acidalia Boeber, 1809

SPEC Boloria selene (Denis & Schiffermüller, 1775)

ORIG Papilio selene Denis & Schiffermüller, 1775

SUBS Boloria selene selene (Denis & Schiffermüller, 1775)
SUBS Boloria selene thalia Hübner, 1790
SUBS Boloria selene dilutior Fixsen, 1887
SUBS Boloria selene chibiana Matsumura, 1927
SUBS Boloria selene perryi Butler, 1882
SUBS Boloria selene myrina (Cramer, 1777)
SUBS Boloria selene nebraskensis (Holland, 1928)
SUBS Boloria selene sabulicollis (Kohler, 1977)
SUBS Boloria selene tolladensis (Barnes & Benjamin, 1925)
SUBS Boloria selene albequina (Holland, 1928)
SUBS Boloria selene atrocostalis (Huard, 1927)
SUBS Boloria selene terraenovae (Holland, 1928)

SPEC Boloria erubescens (Staudinger, 1901)
SPEC Boloria selenis (Eversmann, 1837)

SUBS Boloria selenis selenis (Eversmann, 1837)
SUBS Boloria selenis sibirica Ershov, 1870

SPEC Boloria hegemone (Staudinger, 1881)
SPEC Boloria angarensis (Erschoff, 1870)

SUBS Boloria angarensis angarensis (Erschoff, 1870)
SUBS Boloria angarensis samkoi Sheljuzhko, 1931
SUBS Boloria angarensis alticoa Sushkin & Tshetverikov, 107
SUBS Boloria angarensis herzi Wnukowsky, 1927
SUBS Boloria angarensis miakei Matsumura, 1919

SPEC Boloria oscarus (Eversmann, 1844)

SUBS Boloria oscarus oscarus (Eversmann, 1844)
SUBS Boloria oscarus australis Graeser, 1888

SPEC Boloria freija (Thunberg, 1791)

ORIG Papilio freija Thunberg, 1791

SUBS Boloria freija freija (Thunberg, 1791)
SUBS Boloria freija jakutensis Wnukowsky, 1927
SUBS Boloria freija pallida Elwes, 1899
SUBS Boloria freija tarquinius (Curtis, 1835)
SUBS Boloria freija natazhati (Gibson, 1920)
SUBS Boloria freija nabokovi (D. Stallings & Turner, 1947)
SUBS Boloria freija browni (Higgins, 1953)
SUBS Boloria freija pallida Elwes, 1899

SPEC Boloria polaris (Boisduval, 1828)

SUBS Boloria polaris polaris (Boisduval, 1828)
SUBS Boloria polaris stellata (Masters, 1972)

SPEC Boloria thore (Hübner, 1803)

SUBS Boloria thore thore (Hübner, 1803)
SUBS Boloria thore borealis (Staudinger, 1861)
SUBS Boloria thore transuralensis Sheljuzhko, 1931
SUBS Boloria thore herculea Fruhstorfer, 1907
SUBS Boloria thore hyperusia Fruhstorfer, 1907
SUBS Boloria thore meridionalis Kardakov, 1928

SPEC Boloria hakutozana (Matsumura, 1927)
SPEC Boloria natazhati Gibson, 1920
SPEC Boloria frigga (Thunberg, 1791)

ORIG Papilio frigga Thunberg, 1791

SUBS Boloria frigga frigga (Thunberg, 1791)
SUBS Boloria frigga alpestris Elwes, 1899
SUBS Boloria frigga maritima Kardakov, 1928
SUBS Boloria frigga saga (Staudinger, 1861)
SUBS Boloria frigga alaskensis (Lehmann, 1913)
SUBS Boloria frigga sagata (Barnes & Benjamin, 1923)

SPEC Boloria dia (Linnaeus, 1767)

ORIG Papilio dia Linnaeus, 1767

SUBS Boloria dia dia (Linnaeus, 1767)
SUBS Boloria dia alpina Elwes, 1899
SUBS Boloria dia setania Fruhstorfer, 1909

SPEC Boloria gong (Oberthür, 1884)

SUBS Boloria gong gong (Oberthür, 1884)
SUBS Boloria gong splendens Murayama
SUBS Boloria gong xizangensis Huang, 2000

SPEC Boloria jerdoni (Lang, 1868)

ORIG Argynnis jerdoni Lang, 1868

SPEC Boloria improba (Butler, 1877)

SUBS Boloria improba improba (Butler, 1877)
SUBS Boloria improba improbula (Bryk, 1920)
SUBS Boloria improba youngi (Holland, 1900)
SUBS Boloria improba acrocnema

SPEC Boloria titania (Esper, 1793)

ORIG Papilio titania Esper, 1793
SSYN Papilio amathusia Esper, 1784

SUBS Boloria titania titania (Esper, 1793)
SUBS Boloria titania bivina Fruhstorfer, 1909
SUBS Boloria titania staudingeri Wnukowsky, 1929
SUBS Boloria titania miyakei Matsumura, 1919
SUBS Boloria titania boisduvalii (Duponchel, 1832)
SUBS Boloria titania rainieri (Barnes & McDunnough, 1913)
SUBS Boloria titania grandis (Barnes & McDunnough, 1916)
SUBS Boloria titania montina (Scudder, 1863)
SUBS Boloria titania ingens (Barnes & McDunnough, 1918)
SUBS Boloria titania helena (Edwards, 1871)

SPEC Boloria euphrosyne (Linnaeus, 1758)

ORIG Papilio euphrosyne Linnaeus, 1758

SUBS Boloria euphrosyne euphrosyne (Linnaeus, 1758)
SUBS Boloria euphrosyne nephele Herrich-Schaeffer, 1846
SUBS Boloria euphrosyne oprhanus Fruhstorfer, 1907
SUBS Boloria euphrosyne kamtschadalus Seitz, 1908

SPEC Boloria chariclea (Schneider, 1794)

SUBS Boloria chariclea chariclea (Schneider, 1794)
SUBS Boloria chariclea arctica (Zetterstedt, 1839)
SUBS Boloria chariclea butleri (Edwards, 1883)

SPEC Boloria bellona (Fabricius, 1775)

ORIG Papilio bellona Fabricius, 1775

SUBS Boloria bellona bellona (Fabricius, 1775)
SUBS Boloria bellona toddi (Holland, 1928)
SUBS Boloria bellona jenistae (D. Stallings & Turner, 1947)

SPEC Boloria kriemhild (Strecker, 1879)
SPEC Boloria epithore (Edwards, 1864)

SUBS Boloria epithore epithore (Edwards, 1864)
SUBS Boloria epithore chermocki (E. & S. Perkins, 1966)
SUBS Boloria epithore borealis (E. Perkins, 1973)
SUBS Boloria epithore sierra (E. Perkins, 1973)

SPEC Boloria alberta (Edwards, 1890)

SUBS Boloria alberta alberta (Edwards, 1890)
SUBS Boloria alberta kurentzovi Wyatt, 1961

SPEC Boloria astarte (Doubleday & Hewitson, 1847)
SPEC Boloria tritonia (de Boeber, 1812)
SPEC Boloria distincta (Gibson, 1920)

SUBS Boloria distincta distincta (Gibson, 1920)
SUBS Boloria distincta machati Korshunov, 1987
SUBS Boloria distincta dubatolovi Korshunov, 1987
SUBS Boloria distincta tschukotkensis Wyatt, 1961

SPEC Boloria matveevi Gorbunov & Korshunov, 1995
SPEC Boloria erda (Christoph, 1892)

SUBS Boloria erda erda (Christoph, 1892)
SUBS Boloria erda dulkeiti Kurenzov, 1970

SPEC Boloria elatus (Staudinger, 1892)
SPEC Boloria iphigenia (Graeser, 1888)

SUBS Boloria iphigenia iphigenia (Graeser, 1888)
SUBS Boloria iphigenia sachalinensis Matsumura, 1925

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