Incertae sedis Nymphalinae

Niklas Wahlberg

Updated 2022-08-12. Based on

• Williams, M. C. 2018: Afrotropical Butterflies 17th Edition.
• Markku Savela: Lepidoptera and other lifeforms
• The Butterflies of America
• Zhang, J., Cong, Q., Shen, J., Opler, P. A. and Grishin, N. V. 2021: Genomics-guided refinement of butterfly taxonomy. The Taxonomic Report 9-3.

GENU Pycina Doubleday, [1849]

SPEC Pycina zamba Doubleday, [1849]

SUBS Pycina zamba zamba Doubleday, [1849]
SUBS Pycina zamba zelys Godman & Salvin, 1884

GENU Rhinopalpa Felder & Felder, 1860

SPEC Rhinopalpa polynice (Cramer, 1779)

ORIG Papilio polynice Cramer, 1779

SUBS Rhinopalpa polynice polynice (Cramer, 1779)
SUBS Rhinopalpa polynice eudoxia (Guérin-Méneville, 1840)

ORIG Vanessa eudoxia Guérin-Méneville, 1840
SSYN Rhinopalpa fulva Felder & Felder, 1860

SUBS Rhinopalpa polynice birmana Fruhstorfer, 1898
SUBS Rhinopalpa polynice acharis Fruhstorfer, 1913
SUBS Rhinopalpa polynice helionice Fruhstorfer, 1913
SUBS Rhinopalpa polynice callonice Frushtorfer, 1898
SUBS Rhinopalpa polynice mentawica Frushtorfer, 1913
SUBS Rhinopalpa polynice elpenice (Felder & Felder, 1867)
SUBS Rhinopalpa polynice stratonice (Felder & Felder, 1867)
SUBS Rhinopalpa polynice megalonice (Felder & Felder, 1867)

GENU Kallimoides Shirôzu & Nakanishi, 1984

SPEC Kallimoides rumia (Doubleday, [1849])

ORIG Kallima rumia Doubleday, [1849]

SUBS Kallimoides rumia rumia (Doubleday, [1849])
SUBS Kallimoides rumia jadyae (Fox, 1968)
SUBS Kallimoides rumia rattrayi (Sharpe, 1904)

GENU Vanessula Dewitz, 1887

SPEC Vanessula milca (Hewitson, 1873)

ORIG Liptena milca Hewitson, 1873

SUBS Vanessula milca milca (Hewitson, 1873)
SUBS Vanessula milca angustifasciata Joicey & Talbot, 1928
SUBS Vanessula milca buchneri Dewitz, 1887
SUBS Vanessula milca latifasciata Joicey & Talbot, 1928

GENU Doleschallia Felder & Felder, 1860

GSYN Apatura Hübner, 1819

SPEC Doleschallia bisaltide (Cramer, 1777)

ORIG Papilio bisaltide Cramer, 1777

SUBS Doleschallia bisaltide bisaltide (Cramer, 1777)
SUBS Doleschallia bisaltide indica Moore, 1899
SUBS Doleschallia bisaltide malabarica Fruhstorfer, 1899
SUBS Doleschallia bisaltide ceylonica Fruhstorfer, 1903
SUBS Doleschallia bisaltide mariae Fruhstorfer, 1903
SUBS Doleschallia bisaltide siamensis Fruhstorfer, 1912
SUBS Doleschallia bisaltide philippensis Fruhstorfer, 1899
SUBS Doleschallia bisaltide continentalis Fruhstorfer, 1899
SUBS Doleschallia bisaltide pratipa Felder & Felder, 1860
SUBS Doleschallia bisaltide australis Felder & Felder, 1867
SUBS Doleschallia bisaltide sciron Godman & Salvin, 1888
SUBS Doleschallia bisaltide tualensis Fruhstorfer, 1899
SUBS Doleschallia bisaltide nasica Fruhstorfer, 1907
SUBS Doleschallia bisaltide scapus Fruhstorfer, 1912
SUBS Doleschallia bisaltide apameia Fruhstorfer, 1912
SUBS Doleschallia bisaltide cethega Fruhstorfer, 1912
SUBS Doleschallia bisaltide nigromarginata Joicey & Noakes, 1915
SUBS Doleschallia bisaltide tenimberensis Fruhstorfer, 1912
SUBS Doleschallia bisaltide herrichi Butler, 1876
SUBS Doleschallia bisaltide montrouzieri Butler, 1876
SUBS Doleschallia bisaltide gurelca Grose-Smith & Kirby, 1893
SUBS Doleschallia bisaltide menexema Fruhstorfer, 1912
SUBS Doleschallia bisaltide rennellensis Howarth, 1962
SUBS Doleschallia bisaltide bougainvillensis Strand, 1920
SUBS Doleschallia bisaltide merguiana Evans, 1924
SUBS Doleschallia bisaltide karabachica Tytler, 1940
SUBS Doleschallia bisaltide denisi Viette, 1950
SUBS Doleschallia bisaltide koshunensis Murayama & Shimonoya, 1963

SPEC Doleschallia polibete (Cramer, 1779)

SUBS Doleschallia polibete polibete (Cramer, 1779)
SUBS Doleschallia polibete celebensis Fruhstorfer, 1899
SUBS Doleschallia polibete sulaensis Fruhstorfer, 1899
SUBS Doleschallia polibete maturitas Tsukada, 1985

SPEC Doleschallia tongana Hopkins, 1927

SUBS Doleschallia tongana tongana Hopkins, 1927
SUBS Doleschallia tongana vomana Fruhstorfer, 1902

SPEC Doleschallia nacar (Boisduval, 1832)

SUBS Doleschallia nacar nacar (Boisduval, 1832)
SUBS Doleschallia nacar trachelus Fruhstorfer, 1907

SPEC Doleschallia browni Salvin & Godman, 1877

SUBS Doleschallia browni browni Salvin & Godman, 1877
SUBS Doleschallia browni scotina Fruhstorfer, 1912

SPEC Doleschallia noorna Grose-Smith & Kirby, 1893

SUBS Doleschallia noorna noorna Grose-Smith & Kirby, 1893
SUBS Doleschallia noorna demetria Fruhstorfer, 1912
SUBS Doleschallia noorna fulva Joicey & Noakes, 1915
SUBS Doleschallia noorna lyncurion Fruhstorfer, 1912
SUBS Doleschallia noorna antimia Fruhstorfer, 1912

SPEC Doleschallia dascon Godman & Salvin, 1880

SUBS Doleschallia dascon dascon Godman & Salvin, 1880
SUBS Doleschallia dascon dasconides Fruhstorfer, 1903

SPEC Doleschallia rickardi Grose-Smith, 1890

SUBS Doleschallia rickardi rickardi Grose-Smith, 1890
SUBS Doleschallia rickardi pfeili Honrath, 1892

SPEC Doleschallia melana Staudinger, 1886
SPEC Doleschallia comrii Godman & Salvin, 1878
SPEC Doleschallia dascylus Godman & Salvin, 1880

SUBS Doleschallia dascylus dascylus Godman & Salvin, 1880
SUBS Doleschallia dascylus phalinus Fruhstorfer, 1907
SUBS Doleschallia dascylus anicetus Fruhstorfer, 1915
SUBS Doleschallia dascylus eudascylus Fruhstorfer, 1903
SUBS Doleschallia dascylus olivacea Fruhstorfer, 1912

SPEC Doleschallia hexophthalmos (Gmelin, 1790)

SUBS Doleschallia hexophthalmos hexophthalmos (Gmelin, 1790)
SUBS Doleschallia hexophthalmos gaius Fruhstorfer, 1912
SUBS Doleschallia hexophthalmos solus Fruhstorfer, 1912
SUBS Doleschallia hexophthalmos areus Fruhstorfer, 1907
SUBS Doleschallia hexophthalmos kapaurensis Fruhstorfer, 1899
SUBS Doleschallia hexophthalmos varus Fruhstorfer, 1912
SUBS Doleschallia hexophthalmos donus Fruhstorfer, 1915

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