Junoniini
Updated 2021-08-20. Based on
- Williams, M. C. 2018: Afrotropical Butterflies 17th Edition.
- Markku Savela's pages
- Lamas, G. 2004: Atlas of Neotropical Lepidoptera. Checklist: Part 4A, Hesperioidea - Papilionoidea. Scientific Publishers, Gainsville, Florida.
- Mendes, LF, Bivar-de-Sousa, A, Vasconcelos, S, Lopes, LF 2018: On the butterflies of genus Precis Hubner, 1819 known in Angola, with description of a new species (Lepidoptera: Nymphalidae, Nymphalinae). SHILAP-REVISTA DE LEPIDOPTEROLOGIA, 46 (184):653-672
- American Junonia based partly on Cong, Q., Zhang, J., Shen, J., Cao, X., Brévignon, C. & Grishin, N. V. 2020: Speciation in North American Jnonia from a genomic perspective. Systematic Entomology 45: 803-837. DOI: 10.1111/syen.12428 and partly on personal communication with Christian Brévignon.
- Pyrcz, T. W., et al. 2021: Alpha-taxonomy and phylogeny of African Junoniini butterflies based on morphological data, with an emphasis on genitalia, and COI barcode (Lepidoptera: Nymphalidae). Zootaxa DOI: 10.11646/zootaxa.4991.3.1
- Sáfián, Florczyk & Takano 2023: A new species in the genus Precis Hübner, [1819] (Lepidoptera: Nymphalidae: Nymphalinae), another surprising discovery in the Nimba Mountains, Guinea and the broader Guinea Highlands in West Africa. Zootaxa DOI: 10.11646/zootaxa.5249.4.4
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GENU Junonia Hübner, 1819
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GSYN Alcyoneis Hübner, 1819
GSYN Aresta Billberg, 1820
GSYN Kamilla Collins & Larsen, 1991
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SPEC Junonia genoveva (Cramer, 1780)
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SUBS Junonia genoveva genoveva (Cramer, 1779)
SUBS Junonia genoveva vivida W.T.M. Forbes, [1929]
SUBS Junonia genoveva constricta C. Felder & R. Felder, 1867
SUBS Junonia genoveva incarnata C. Felder & R. Felder, 1867
SUBS Junonia genoveva infuscata C. Felder & R. Felder, 1867
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SSYN Junonia lavinia var. basifusca Weymer, 1890
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SSYN Papilio cadmus Larrañaga, 1923
SPEC Junonia neildi Brévignon, 2004
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SUBS Junonia neildi neildi Brévignon, 2004
SUBS Junonia neildi varia Grishin, 2020
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SUBS Junonia zonalis zonalis C. Felder & R. Felder, 1867
SUBS Junonia zonalis swifti Brévignon, 2004
SUBS Junonia zonalis michaelisi Fruhstorfer, 1907
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SUBS Junonia coenia coenia Hübner, [1822]
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SSYN Junonia coenia weidenhammeri Polacek, 1925
SSYN Precis coena [sic] f. rubrosuffusa Field, 1936
SSYN Junonia coenia f. rosa Whittaker & Stallings, 1944
SSYN Papilio quinatzin Arias, 1968
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SSYN Junonia coenia ab. schraderi Gunder, 1925
SSYN Junonia coenia tr. f. wilhelmi Gunder, 1927
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SSYN Junonia nigrosuffusa var melanina A. Hall, 1919
SPEC Junonia pacoma Grishin, 2020
SPEC Junonia wahlbergi Brévignon, 2008
SPEC Junonia flirtea (Fabricius, 1793)
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SSYN Junonia hübneri [sic] W.F. Kirby, [1900]
SSYN Precis lavinia ab. obscurata Kivirikko, 1936
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SUBS Junonia evarete evarete (Cramer, 1779)
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OSYN Papilio lavinia Cramer, 1775
OSYN Papilio cortes Herbst, 1796
SSYN Papilio esra Fabricius, 1798
SUBS Junonia evarete dougueti Brévignon, 2008
SUBS Junonia evarete oscura Neild, 2008
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SSYN Junonia evarete ab nigralis WTM Forbes, [1929]
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SSYN Precis lavinia funebris Bryk, 1953
SUBS Junonia evarete occidentalis C. Felder & R. Felder, 1862
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SSYN Junonia negra C. Felder & R. Felder, 1867
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SUBS Junonia divaricata divaricata C. Felder & R. Felder, 1867
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SSYN Junonia divaricata arenosa W.T.M. Forbes, [1929]
SUBS Junonia divaricata mitaraka Brévignon, 2012
SPEC Junonia vestina C. Felder & R. Felder, 1867
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SUBS Junonia vestina vestina C. Felder & R. Felder, 1867
SUBS Junonia vestina livia Fruhstorfer, 1912
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ORIG Papilio orithya Linnaeus, 1758
SSYN Precis orithya (Linnaeus, 1758)
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SUBS Junonia orithya orithya (Linnaeus, 1758)
SUBS Junonia orithya patenas (Fruhstorfer, 1912)
SUBS Junonia orithya ocyale Hübner, 1819
SUBS Junonia orithya hainanensis (Fruhstorfer, 1912)
SUBS Junonia orithya wallacei Distant, 1883
SUBS Junonia orithya swinhoei Butler, 1885
SUBS Junonia orithya minagara (Fruhstorfer, 1904)
SUBS Junonia orithya sumatrana Fruhstorfer, 1906
SUBS Junonia orithya baweana Fruhstorfer, 1906
SUBS Junonia orithya mevaria (Fruhstorfer, 1904)
SUBS Junonia orithya metion Fruhstorfer, 1905
SUBS Junonia orithya leucasia (Fruhstorfer, 1912)
SUBS Junonia orithya celebensis Staudinger, 1888
SUBS Junonia orithya novaguineae Hagen, 1897
SUBS Junonia orithya orthosia (Godart, 1824)
SUBS Junonia orithya eutychia (Fruhstorfer, 1912)
SUBS Junonia orithya palea (Fruhstorfer, 1912)
SUBS Junonia orithya neopommerana Ribbe, 1898
SUBS Junonia orithya albicincta Butler, 1875
SUBS Junonia orithya madagascariensis Guenée, 1865
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SSYN Junonia booepis Trimen, 1879
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SSYN Precis orithya cheesmani Riley, 1925
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SUBS Junonia oenone oenone (Linnaeus, 1758)
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SSYN Papilio clelia Cramer, 1775
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ORIG Papilio villida Fabricius, 1787
SSYN Vanessa calybe Godart, 1819
SSYN Precis villida (Fabricius, 1787)
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ORIG Papilio hierta Fabricius, 1798
SSYN Papilio lintingensis Osbeck, 1765
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SUBS Junonia hierta hierta (Fabricius, 1798)
SUBS Junonia hierta cebrene Trimen, 1870
SUBS Junonia hierta paris Trimen, 1887
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SSYN Junonia ixia Butler, 1866
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SUBS Junonia westermanni westermanni Westwood, 1870
SUBS Junonia westermanni suffusa (Rothschild & Jordan, 1903)
SUBS Junonia westermanni splendens (Schmidt, 1921)
SPEC Junonia artaxia (Hewitson, 1864)
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ORIG Precis artaxia Hewitson, 1864
SSYN Precis nachtigali Dewitz, 1879
SSYN Precis nobilitata Thurau, 1903
SPEC Junonia iphita (Cramer, 1779)
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ORIG Papilio iphita Cramer, 1779
SSYN Precis iphita (Cramer, 1779)
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SUBS Junonia iphita iphita (Cramer, 1779)
SUBS Junonia iphita pluviatilis Fruhstorfer
SUBS Junonia iphita horsfieldi Moore, 1899
SUBS Junonia iphita tosca Fruhstorfer
SUBS Junonia iphita viridis Staudinger
SUBS Junonia iphita adelaida Staudinger
SUBS Junonia iphita cebara Fruhstorfer
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ORIG Papilio hedonia Linnaeus, 1764
SSYN Precis hedonia (Linnaeus, 1764)
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SUBS Junonia hedonia hedonia (Linnaeus, 1764)
SUBS Junonia hedonia ida (Cramer, 1775)
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ORIG Papilio ida Cramer, 1775
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ORIG Precis seitzi Corbet, 1937
SUBS Junonia hedonia hondai Hayashi
SUBS Junonia hedonia hellanis Felder
SUBS Junonia hedonia numana Fruhstorfer
SUBS Junonia hedonia thero Fruhstorfer
SUBS Junonia hedonia zelima (Fabricius, 1775)
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ORIG Papilio zelima Fabricius, 1775
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SSYN Junonia shmiedeli Fiedler, 1920
SPEC Junonia sophia (Fabricius, 1793)
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ORIG Papilio sophia Fabricius, 1793
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SUBS Junonia sophia sophia (Fabricius, 1793)
SUBS Junonia sophia infracta Butler, 1888
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SSYN Precis sophia albida Suffert, 1904
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SUBS Junonia terea terea (Drury, 1773)
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SSYN Junonia zipha Butler, 1869
SUBS Junonia terea fumata (Rothschild & Jordan, 1903)
SPEC Junonia goudoti (Boisduval, 1833)
SPEC Junonia natalica (Felder & Felder, 1860)
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SUBS Junonia natalica natalica (Felder & Felder, 1860)
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SSYN Junonia hecate Trimen, 1862
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SSYN Vanessa orthosia Klug, 1845
SSYN Salamis ethyra Feisthamel, 1850
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ORIG Papilio atlites Linnaeus, 1763
SSYN Papilio laomedia Linnaeus, 1767
SSYN Precis atlites (Linnaeus, 1763)
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ORIG Papilio almana Linnaeus, 1758
SSYN Alcyoneis almane Hübner, 1819
SSYN Precis almana (Linnaeus, 1758)
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SUBS Junonia almana almana (Linnaeus, 1758)
SUBS Junonia almana sumbae Doherty
SUBS Junonia almana battana Fruhstorfer
SUBS Junonia almana javana Felder, 1862
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ORIG Papilio lemonias Linnaeus, 1758
SSYN Precis lemonias (Linnaeus, 1758)
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SUBS Junonia lemonias lemonias (Linnaeus, 1758)
SUBS Junonia lemonias vaisya Fruhstorfer
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ORIG Papilio erigone Cramer, 1775
SSYN Precis walkeri Butler, 1901
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SUBS Junonia erigone erigone (Cramer, 1775)
SUBS Junonia erigone iona Grose-Smith
SUBS Junonia erigone tristis Miskin
SUBS Junonia erigone leucophora Fruhstorfer
SUBS Junonia erigone expansa Butler
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SSYN Precis timorensis (Wallace, 1869)
SPEC Junonia intermedia (Felder & Felder, 1867)
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ORIG Precis intermedia Felder & Felder, 1867
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SUBS Junonia intermedia intermedia (Felder & Felder, 1867)
SUBS Junonia intermedia teurnia Fruhstorfer
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SSYN Junonia pavonina Butler, 1895
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SSYN Kallima incerta Grünberg, 1908
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SUBS Junonia cymodoce cymodoce (Cramer, 1777)
SUBS Junonia cymodoce lugens (Schultze, 1912)
SPEC Junonia cytora (Doubleday, 1847)
SPEC Junonia temora (Felder & Felder, 1867)
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SUBS Junonia temora temora (Felder & Felder, 1867)
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SSYN Salamis amarantha Butler, 1869
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SPEC Salamis anteva (Ward, 1870)
SPEC Salamis augustina Boisduval, 1833
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SUBS Salamis augustina augustina Boisduval, 1833
SUBS Salamis augustina vinsoni Le Cerf, 1922
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SSYN Salamis cacta languida Bartel, 1905
SSYN Salamis strandi Röber, 1937
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SUBS Salamis amaniensis amaniensis Vosseler, 1907
SUBS Salamis amaniensis eileenae Henning & Joannou, 1994
SUBS Salamis amaniensis humbloti Turlin, 1994
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SPEC Yoma sabina (Cramer, 1780)
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ORIG Papilio sabina Cramer, 1780
SSYN Salamis australis Fruhstorfer, 1899
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SUBS Yoma sabina sabina (Cramer, 1780)
SUBS Yoma sabina parva (Butler, 1876)
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ORIG Rhinopalpa parva Butler, 1876
SUBS Yoma sabina atomaria Fruhstorfer
SUBS Yoma sabina vasuki Doherty, 1886
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SUBS Yoma algina algina (Boisduval, 1832)
SUBS Yoma algina netonia Fruhstorfer
SUBS Yoma algina vestina Fruhstorfer
SUBS Yoma algina manusi Rothschild
SUBS Yoma algina odilla Fruhstorfer
SUBS Yoma algina kokopona Hagen
SUBS Yoma algina helisson Fruhstorfer
SUBS Yoma algina pavonia Mathew
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SPEC Protogoniomorpha anacardii (Linnaeus, 1758)
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ORIG Papilio anacardii Linnaeus, 1758
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SUBS Protogoniomorpha anacardii anacardii (Linnaeus, 1758)
SUBS Protogoniomorpha anacardii ansorgei (Rothschild, 1904)
SPEC Protogoniomorpha duprei (Vinson, 1863)
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SSYN Salamis definata Butler, 1879
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SSYN Papilio aethiops Palisot de Beauvais, 1805
SSYN Vanessa aglatonice Godart, 1819
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GSYN Coryphaeola Butler, 1878
GSYN Kallimula Holland, 1920
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SPEC Precis octavia (Cramer, 1777)
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ORIG Papilio octavia Cramer, 1777
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SUBS Precis octavia octavia (Cramer, 1777)
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SSYN Papilio amestris Drury, 1782
SSYN Papilio zingha Fabricius, 1787
SSYN Vanessa emma Godart, 1819
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SSYN Junonia calescens Butler, 1894
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ORIG Salamis ceryne Boisduval, 1847
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SUBS Precis ceryne ceryne (Boisduval, 1847)
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SSYN Salamis tukuoa Wallengren, 1857
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SSYN Junonia trimeni Butler, 1894
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SSYN Precis simia Wallengren, 1857
SSYN Precis ibris Felder & Felder, 1867
SSYN Junonia micromera Butler, 1876
SSYN Precis petersi Dewitz, 1879
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SSYN Vanessa musa Guérin-Méneville, 1844
SSYN Precis boisduvali Staudinger, 1885
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SSYN Papilio leodice Cramer, 1777
SSYN Papilio harpyia Fabricius, 1781
SSYN Papilio trullus Herbst, 1794
SSYN Vanessa galami Boisduval, 1833
SSYN Precis monroviana Staudinger, 1885
SPEC Precis actia Distant, 1880
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SSYN Precis vetula Staudinger, 1885
SSYN Precis pelarga albofasciata Suffert, 1904
SSYN Precis actia rubrofasciata Suffert, 1904
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SUBS Precis sinuata sinuata Plötz, 1880
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SSYN Precis serena Weymer, 1892
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SSYN Kallimula osborni Holland, 1920
SSYN Precis rauana kakamega Carcasson, 1961
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SSYN Precis omissa Rothschild, 1918
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SUBS Precis tugela tugela Trimen, 1879
SUBS Precis tugela aurorina (Butler, 1894)
SUBS Precis tugela pyriformis (Butler, 1896)
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SUBS Precis milonia milonia Felder & Felder, 1867
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SSYN Junonia kowara Ward, 1871
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ORIG Kallima eurodoce Westwood, 1850
SPEC Precis archesia (Cramer, 1779)
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SUBS Precis archesia archesia (Cramer, 1779)
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SSYN Vanessa pelasgis Godart, 1819
SSYN Junonia chapunga Hewitson, 1864
SSYN Precis staudingeri Dewitz, 1879
SSYN Precis guruana Rogenhofer, 1891
SPEC Precis limnoria (Klug, 1845)
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SUBS Precis limnoria limnoria (Klug, 1845)
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SSYN Precis limnoria niveisticta Gabriel, 1954
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GSYN Esoptria Hübner, 1819
GSYN Diadema Boisduval, 1832 (preocc. Diadema Gray, 1825)
GSYN Euralia Westwood, 1850
GSYN Eucalia Felder, 1861
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SPEC Hypolimnas alimena (Linnaeus, 1758)
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ORIG Papilio alimena Linnaeus, 1758
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SUBS Hypolimnas alimena alimena (Linnaeus, 1758)
SUBS Hypolimnas alimena heteromorpha Röber, 1891
SUBS Hypolimnas alimena senia Fruhstorfer, 1912
SUBS Hypolimnas alimena bandana Fruhstorfer, 1912
SUBS Hypolimnas alimena remigia Fruhstorfer, 1912
SUBS Hypolimnas alimena eligia Fruhstorfer, 1912
SUBS Hypolimnas alimena saturnia Fruhstorfer, 1903
SUBS Hypolimnas alimena polymena (Felder & Felder, 1867)
SUBS Hypolimnas alimena forbesi Butler, 1883
SUBS Hypolimnas alimena eremita Butler, 1883
SUBS Hypolimnas alimena curicta Fruhstorfer, 1912
SUBS Hypolimnas alimena libisonia Fruhstorfer, 1912
SUBS Hypolimnas alimena bateia Fruhstorfer, 1915
SUBS Hypolimnas alimena afra Fruhstorfer, 1903
SUBS Hypolimnas alimena obsolescens Fruhstorfer, 1903
SUBS Hypolimnas alimena lamina Fruhstorfer, 1903
SUBS Hypolimnas alimena darwinensis Waterhouse & Lyell, 1914
SUBS Hypolimnas alimena inexpectata (Godman & Salvin, 1877)
SUBS Hypolimnas alimena salomonis Ribbe, 1898
SUBS Hypolimnas alimena ysabela Fruhstorfer, 1912
SUBS Hypolimnas alimena fulginescens (Mathew, 1887)
SUBS Hypolimnas alimena diffusa Howarth, 1962
SUBS Hypolimnas alimena libateia Howarth, 1962
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ORIG Diadema anomala Wallace, 1869
SSYN Diadema albula Wallace, 1869
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SUBS Hypolimnas anomala anomala (Wallace, 1869)
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ORIG Hypolimnas nivas Fruhstorfer, 1913
SUBS Hypolimnas anomala arnoldi Fruhstorfer, 1903
SUBS Hypolimnas anomala interstincta (Butler, 1873)
SUBS Hypolimnas anomala wallaceana (Butler, 1873)
SUBS Hypolimnas anomala euvaristos Fruhstorfer, 1912
SUBS Hypolimnas anomala truentus Fruhstorfer, 1912
SPEC Hypolimnas anthedon (Doubleday, 1845)
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SUBS Hypolimnas anthedon anthedon (Doubleday, 1845)
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SSYN Papilio dubius Palisot de Beauvois, 1805 (preocc.)
SSYN Diadema damoclina Trimen, 1869
SSYN Hypolimnas daemona Staudinger, 1896
SSYN Hypolimnas dubius poensis Rothschild, 1918
SSYN Hypolimnas macarthuri Neidhoefer, 1972
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SSYN Diadema mima Trimen, 1869
SSYN Hypolimnas angustilimbata Weymer, 1892
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SSYN Panopea bewsheri Butler, 1879
SSYN Panopea diffusa Butler, 1880
SSYN Hypolimnas madagascariensis Mabille, 1881
SSYN Hypolimnas lutescens Mabille, 1887
SSYN Hypolimnas limbata Crowley, 1890
SSYN Hypolimnas dubius mayottensis Rothschild, 1918
SSYN Hypolimnas dubia antelmei Le Cerf, 1928
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ORIG Papilio antilope Cramer, 1777
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SUBS Hypolimnas antilope antilope (Cramer, 1777)
SUBS Hypolimnas antilope mela Fruhstorfer, 1903
SUBS Hypolimnas antilope maglovius Fruhstorfer, 1912
SUBS Hypolimnas antilope quinctinus Fruhstorfer, 1912
SUBS Hypolimnas antilope shortlandia Ribbe, 1898
SUBS Hypolimnas antilope wagneri Clark, 1946
SUBS Hypolimnas antilope scopas (Godman & Salvin, 1888)
SUBS Hypolimnas antilope albomela Howarth, 1962
SPEC Hypolimnas bartelotti Grose-Smith, 1890
SPEC Hypolimnas bolina (Linnaeus, 1758)
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ORIG Papilio bolina Linnaeus, 1758
SSYN Hypolimnas parva Aurivillius, 1920
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SUBS Hypolimnas bolina bolina (Linnaeus, 1758)
SUBS Hypolimnas bolina nerina (Fabricius, 1775)
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SSYN Papilio iphigenia Cramer, 1775
SSYN Papilio alcmene Cramer, 1775
SSYN Papilo perimele Cramer, 1775
SSYN Papilio proserpina Cramer, 1779
SSYN Hypolimnas pseudomisippus Fruhstorfer, 1912
SUBS Hypolimnas bolina pulchra (Butler, 1874)
SUBS Hypolimnas bolina pallescens (Butler, 1874)
SUBS Hypolimnas bolina lisianassa (Cramer, 1779)
SUBS Hypolimnas bolina jacintha (Drury, 1773)
SUBS Hypolimnas bolina incommoda Butler, 1879
SUBS Hypolimnas bolina kezia (Butler, 1877)
SPEC Hypolimnas deceptor (Trimen, 1873)
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ORIG Diadema deceptor Trimen, 1873
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SUBS Hypolimnas deceptor deceptor (Trimen, 1873)
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SSYN Euralia kirbyi Butler, 1898
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SSYN Euralia grandidieri Mabille, 1899
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SUBS Hypolimnas deois deois (Hewitson, 1858)
SUBS Hypolimnas deois divina Fruhstorfer, 1903
SUBS Hypolimnas deois tydea (Felder & Felder, 1867)
SUBS Hypolimnas deois obianus Fruhstorfer, 1912
SUBS Hypolimnas deois hewitsoni (Wallace, 1869)
SUBS Hypolimnas deois panopion Grose-Smith, 1894
SUBS Hypolimnas deois denticulata Talbot, 1932
SUBS Hypolimnas deois palladius Grose-Smith, 1897
SUBS Hypolimnas deois paleutes Grose-Smith, 1897
SUBS Hypolimnas deois woodlarkiana Talbot, 1932
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SSYN Diadema imperialis Guenée, 1865
SPEC Hypolimnas dinarcha (Hewitson, 1865)
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SUBS Hypolimnas dinarcha dinarcha (Hewitson, 1865)
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SSYN Hypolimnas dinarcha narchadi Suffert, 1904
SSYN Hypolimnas dinarcha liberiensis Bernardi, 1959
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SUBS Hypolimnas diomea diomea Hewitson, 1861
SUBS Hypolimnas diomea fraterna (Wallace, 1869)
SPEC Hypolimnas euploeoides Rothschild, 1915
SPEC Hypolimnas exiguus Samson, 1980
SPEC Hypolimnas inopinata Waterhouse, 1920
SPEC Hypolimnas mechowi (Dewitz, 1884)
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SSYN Diadema ragiens Capronnier, 1889
SSYN Hypolimnas stanleyi Grose-Smith, 1890
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SSYN Papilio diocippus Cramer, 1775
SSYN Papilio maria Cramer, 1779
SSYN Papilio inaria Cramer, 1779
SSYN Hypolimnas alcippoides Butler, 1883
SSYN Hypolimnas picta Fruhstorfer, 1912
SSYN Hypolimnas immima Bernardi, 1959
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SUBS Hypolimnas monteironis monteironis (Druce, 1874)
SUBS Hypolimnas monteironis major Rothschild, 1918
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SUBS Hypolimnas octocula octocula (Butler, 1869)
SUBS Hypolimnas octocula perryi (Butler, 1876)
SUBS Hypolimnas octocula elsina (Butler, 1877)
SUBS Hypolimnas octocula samoa Samson, 1986
SUBS Hypolimnas octocula futunaensis Samson, 1986
SUBS Hypolimnas octocula tanna Samson, 1986
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ORIG Papilio pandarus Linnaeus, 1758
SSYN Papilio pipleis Linnaeus, 1758
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SUBS Hypolimnas pandarus pandarus (Linnaeus, 1758)
SUBS Hypolimnas pandarus pandora (Wallace, 1869)
SUBS Hypolimnas pandarus hewitsoni (Wallace, 1869)
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SUBS Hypolimnas pithoeka pithoeka Kirsch, 1877
SUBS Hypolimnas pithoeka vulcanica Rothschild, 1915
SUBS Hypolimnas pithoeka fumosus Joicey & Noakes, 1915
SUBS Hypolimnas pithoeka bradleyi Howarth, 1962
SUBS Hypolimnas pithoeka ferruginea Howarth, 1962
SUBS Hypolimnas pithoeka gretheri Clark, 1946
SUBS Hypolimnas pithoeka salomona D'Abrera, 1977
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SUBS Hypolimnas salmacis salmacis (Drury, 1773)
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SSYN Papilio omphale Stoll, 1790
SSYN Hypolimnas salmacis cissalma Suffert, 1904
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SSYN Hypolimnas salmacis ulysses Collins, 1987
SUBS Hypolimnas salmacis platydema Rothschild & Jordan, 1903
SUBS Hypolimnas salmacis thomensis Aurivillius, 1910
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SUBS Hypolimnas saundersii saundersii Hewitson, 1869
SUBS Hypolimnas saundersii junia Fruhstorfer, 1903
SPEC Hypolimnas usambara (Ward, 1872)
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SSYN Hypolimnas imperialis Staudinger, 1885