Euptychiina

Niklas Wahlberg

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Updated 2023-06-17.

Mainly based on:
Espeland, M., et al. 2023: Combining target enrichment and Sanger sequencing data to clarify the systematics of the diverse Neotropical butterfly subtribe Euptychiina (Nymphalidae, Satyrinae). Systematic Entomology doi: 10.1111/syen.12590

Other sources listed at the bottom of this page.

SUBT Euptychiina

GENU Euptychia Hübner, 1818

GSYN Caenoptychia Le Cerf 1919
GSYN Ristia Gagarin 1936

SPEC Euptychia alacristata A. F. E. Neild, Nakahara & S. Fratello, 2014
SPEC Euptychia aquila S. Fratello, Nakahara & Brévignon, 2015
SPEC Euptychia atlantica Nakahara & A.V.L. Freitas, 2017
SPEC Euptychia attenboroughi Neild, Nakahara, S. Fratello & Le Crom, 2015
SPEC Euptychia audacia Brévignon, S. Fratello & Nakahara, 2015
SPEC Euptychia boulleti Le Cerf, 1919

SSYN Euptychia virgata Joicey & Talbot, 1924
SSYN Euptychia tigrina Gagarin, 1936

SPEC Euptychia cesarense Pulido, Andrade, C. Peña & Lamas, 2011

SUBS Euptychia cesarense cesarense Pulido, Andrade, C. Peña & Lamas, 2011
SUBS Euptychia cesarense obtusa Nakahara, 2015
SUBS Euptychia cesarense viloriai Andrade, Pulido, C. Peña & Lamas, 2011

SPEC Euptychia efraini Ríos-Malaver, 2019

HOMO Euptychia similis Henao-Bañol, 2019 (Homonym of Euptychia similis Butler, 1867)

SPEC Euptychia enyita Nakahara, Lamas & Willmott, 2015
SPEC Euptychia enyo Butler, 1867
SPEC Euptychia favonius Nakahara, G. Vega & Willmott, 2016
SPEC Euptychia fernandae Nakahara & Willmott, 2015
SPEC Euptychia fetna Butler, 1870
SPEC Euptychia granatina Nakahara, J.F. LeCrom & J. Hall, 2015
SPEC Euptychia hannemanni Forster, 1964
SPEC Euptychia insolata Butler & H. Druce, 1872

SSYN Euptychia macrophthalma Staudinger, 1876

SPEC Euptychia jesia Butler, 1869
SPEC Euptychia juanjoi J.F. LeCrom, Nakahara & Lamas, 2015
SPEC Euptychia lacandona A. Warren & Nakahara, 2015
SPEC Euptychia marceli Brévignon, 2005

SUBS Euptychia marceli marceli Brévignon, 2005
SUBS Euptychia marceli divisa Benmesbah, Costa, Attal & Viloria, 2021

SPEC Euptychia meta Weymer, 1911
SPEC Euptychia mollina (Hübner, [1813])

SUBS Euptychia mollina mollina (Hübner, [1813])
SUBS Euptychia mollina suzannae Brévignon, 2005

SPEC Euptychia mollis Staudinger, 1876
SPEC Euptychia neblina A. Warren & Nakahara, 2015
SPEC Euptychia neildi Brévignon, 2005
SPEC Euptychia padroni Nakahara, Lamas & Willmott, 2015
SPEC Euptychia pegasus Nakahara & Hall, 2015
SPEC Euptychia picea Butler, 1867
SPEC Euptychia pillaca Nakahara & Willmott, 2015
SPEC Euptychia roraima Nakahara, Fratello & Harvey, 2014
SPEC Euptychia rubrofasciata Miller & Miller, 1988
SPEC Euptychia rufocincta Weymer, 1911
SPEC Euptychia sarah Benmesbah & Viloria, 2019
SPEC Euptychia sophiae Zacca, Nakahara, Dolibaina & Dias, 2015
SPEC Euptychia truncata Nakahara & J. Hall, 2015
SPEC Euptychia westwoodi Butler, 1867

SUBS Euptychia westwoodi westwoodi Butler, 1867
SUBS Euptychia westwoodi muli Brévignon, 2015

SPEC Euptychia woroina Viloria & Benmesbah, 2020

GENU Atlanteuptychia Freitas, Barbosa & Mielke, 2013

SPEC Atlanteuptychia ernestina (Weymer, 1911)

GENU Cyllopsis R. Felder, 1869

SPEC Cyllopsis argentella (Butler & H. Druce, 1872)
SPEC Cyllopsis caballeroi Beutelspacher, 1982
SPEC Cyllopsis clinas (Godman & Salvin, 1889)
SPEC Cyllopsis diazi L. Miller, 1974
SPEC Cyllopsis dospassosi L. Miller, 1974
SPEC Cyllopsis emilia I. Chacón & Nishida, 2002
SPEC Cyllopsis gemma (Hübner, 1808)

SUBS Cyllopsis gemma gemma (Hübner, 1808)
SUBS Cyllopsis gemma freemani (D. Stallings & J. Turner, 1947)

SPEC Cyllopsis guatemalena L. Miller, 1974
SPEC Cyllopsis hedemanni R. Felder, 1869

SUBS Cyllopsis hedemanni hedemanni R. Felder, 1869
SUBS Cyllopsis hedemanni tamaulipensis L. Miller, 1974
SUBS Cyllopsis hedemanni vetones (Godman & Salvin, 1878)

SPEC Cyllopsis hilaria (Godman, 1901)
SPEC Cyllopsis jacquelineae L. Miller, 1974
SPEC Cyllopsis nayarit (R. Chermock, 1947)
SPEC Cyllopsis nelsoni (Godman & Salvin, 1881)
SPEC Cyllopsis pallens L. Miller, 1974
SPEC Cyllopsis parvimaculata L. Miller, 1974
SPEC Cyllopsis pephredo (Godman, 1901)
SPEC Cyllopsis perplexa L. Miller, 1974
SPEC Cyllopsis pertepida (Dyar, 1912)

SUBS Cyllopsis pertepida pertepida (Dyar, 1912)
SUBS Cyllopsis pertepida maniola (Nabokov, 1942)
SUBS Cyllopsis pertepida avicula (Nabokov, 1942)
SUBS Cyllopsis pertepida intermedia L. Miller, 1974

SPEC Cyllopsis philodice (Godman & Salvin, 1878)
SPEC Cyllopsis pseudopephredo (R. Chermock, 1947)
SPEC Cyllopsis pyracmon (Butler, 1867)

SUBS Cyllopsis pyracmon pyracmon (Butler, 1867)
SUBS Cyllopsis pyracmon henshawi (W. H. Edwards, 1876)

SPEC Cyllopsis rogersi (Godman & Salvin, 1878)
SPEC Cyllopsis schausi L. Miller, 1974
SPEC Cyllopsis steinhauserorum L. Miller, 1974
SPEC Cyllopsis suivalenoides L. Miller, 1974
SPEC Cyllopsis suivalens (Dyar, 1914)

SUBS Cyllopsis suivalens suivalens (Dyar, 1914)
SUBS Cyllopsis suivalens escalantei L. Miller, 1974

SPEC Cyllopsis tomemmeli A. Warren & Nakahara, 2018
SPEC Cyllopsis whiteorum L. Miller & J. Maza, 1984
SPEC Cyllopsis wellingi L. Miller, 1978
SPEC Cyllopsis windi L. Miller, 1974

GENU Inbio Nakahara & Espeland, 2015

SPEC Inbio hilara (C. Felder & R. Felder, 1867)

GENU Paramacera Butler, 1868

SPEC Paramacera allyni L. Miller, 1972
SPEC Paramacera xicaque (Reakirt, [1867])
SPEC Paramacera rubrosuffusa L. Miller, 1972
SPEC Paramacera copiosa L. Miller, 1972
SPEC Paramacera chinanteca L. Miller, 1972

GENU Carminda Ebert & Dias, 1998

SPEC Carminda griseldis (Weymer, 1911)
SPEC Carminda paeon (Godart, [1824])
SPEC Carminda umuarama Ebert & Dias, 1997
SPEC Carminda surpresa Barbosa, Aguiar, Rosa, Zacca & Freitas, 2020

GENU Cissia E. Doubleday, 1848

GSYN Argyreuptychia Forster 1964

SPEC Cissia anabelae (L. Miller, 1976)
SPEC Cissia cheneyorum (R. Chermock, 1949)
SPEC Cissia pseudocleophes (L. Miller, 1976)
SPEC Cissia penelope (Fabricius, 1775)

SSYN Euptychia moneta Weymer, 1911

SPEC Cissia pompilia (C. Felder & R. Felder, 1867)

SSYN Euptychia austera Butler, 1867
SSYN Euptychia usitata Butler, 1867
SSYN Euptychia pieria Butler, 1867
SSYN Euptychia thelete Snellen, 1887

SPEC Cissia proba (Weymer, 1911)
SPEC Cissia eous (Butler, 1867)

SSYN Paryphthimoides kiliani Anken, 1999

SPEC Cissia phronius (Godart, [1824])
SPEC Cissia cleophes (Godman & Salvin, 1889)
SPEC Cissia rubricata (W. H. Edwards, 1871)

SUBS Cissia rubricata rubricata (W. H. Edwards, 1871)
SUBS Cissia rubricata smithorum (Wind, 1946)

SPEC Cissia wahala Grishin, 2022

GENU Graphita Nakahara, Marín & Barbosa, 2016

SPEC Graphita griphe (C. Felder & R. Felder, 1867)

GENU Koutalina Viloria & Murienne, 2020

SPEC Koutalina pamela (Hayward, 1957)

GENU Megisto Hübner, 1819

GSYN Palaeonympha Butler, 1871

SPEC Megisto cymela (Cramer, 1777)

SUBS Megisto cymela cymela (Cramer, 1777)
SUBS Megisto cymela viola (Maynard, 1891)

SPEC Megisto opalina (Butler, 1871)

SUBS Megisto opalina opalina (Butler, 1871)
SUBS Megisto opalina macrophthalmia Fruhstorfer, 1911
SUBS Megisto opalina bailiensis Yoshino, 2001

GENU Moneuptychia Forster, 1964

SPEC Moneuptychia castrensis (Schaus, 1902)
SPEC Moneuptychia giffordi Freitas, Emery & Mielke, 2010
SPEC Moneuptychia itapeva Freitas, 2007
SPEC Moneuptychia melchiades (Butler, 1877)
SPEC Moneuptychia montana Freitas, 2015
SPEC Moneuptychia pervagata Freitas, Siewert & Mielke, 2015
SPEC Moneuptychia romanina (Bryk, 1953)
SPEC Moneuptychia soter (Butler, 1877)
SPEC Moneuptychia vitellina Freitas & Barbosa, 2015
SPEC Moneuptychia viviana (Romieux, 1927)
SPEC Moneuptychia wahlbergi Freitas, Barbosa, Siewert & Mielke, 2015

GENU Pharneuptychia Forster, 1964

SPEC Pharneuptychia estoraquensis Henao & Meneses, 2018
SPEC Pharneuptychia haywardiana Lamas, 2010

SSYN Pharneuptychia boliviana (Hayward, 1957)

SPEC Pharneuptychia phares (Godart, [1824])
SPEC Pharneuptychia pharnabazos (Bryk, 1953)
SPEC Pharneuptychia pharnaces (Weymer, 1911)

GENU Stegosatyrus Zacca, Mielke & Pyrcz, 2013

SPEC Stegosatyrus hemiclara Pyrcz, Boyer & Zacca, 2013
SPEC Stegosatyrus imbrialis (Weeks, 1901)
SPEC Stegosatyrus ocelloides (Schaus, 1902)
SPEC Stegosatyrus periphas (Godart, [1824])

GENU Vanima Zacca, Casagrande & Mielke, 2020

SPEC Vanima labe (Butler, 1870)
SPEC Vanima lesbia (Staudinger, [1886])
SPEC Vanima palladia (Butler, 1867)

GENU Yphthimoides Forster, 1964

SPEC Yphthimoides acmenis (Hübner, 1823)
SPEC Yphthimoides angularis (Butler, 1867)
SPEC Yphthimoides bella Barbosa & Freitas, 2015
SPEC Yphthimoides blanquita Barbosa, Marín & Freitas, 2017
SPEC Yphthimoides borasta (Schaus, 1902)
SPEC Yphthimoides celmis (Godart, [1824])
SPEC Yphthimoides cipoensis Freitas, 2004
SPEC Yphthimoides gabriela Barbosa, Freitas & Paluch, 2015
SPEC Yphthimoides inornata (Hayward, 1962)
SPEC Yphthimoides iserhardi Freitas & Barbosa, 2015
SPEC Yphthimoides kinyoni Lamas, Nakahara & Barbosa, 2021
SPEC Yphthimoides leguialimai (Dyar, 1913)
SPEC Yphthimoides manasses (C. Felder & R. Felder, 1867)
SPEC Yphthimoides nareta Barbosa & Freitas, 2017
SPEC Yphthimoides ochracea (Butler, 1867)
SPEC Yphthimoides ordinaria Freitas, Kaminski & Mielke, 2012
SPEC Yphthimoides pacta (Weymer, 1911)
SPEC Yphthimoides patricia (Hayward, 1957)

SSYN Yphthimoides saltuensis (Hayward, 1962)

SPEC Yphthimoides renata (Stoll, 1780)
SPEC Yphthimoides straminea (Butler, 1867)
SPEC Yphthimoides yphthima (C. Felder & R. Felder, 1867)

GENU Cristalinaia Mota, Zacca & Freitas, 2019

SPEC Cristalinaia vitoria Mota, Zacca & Freitas, 2019

GENU Lazulina Willmott, Nakahara & Espeland, 2023

SPEC Lazulina catharina (Staudinger, [1886])
SPEC Lazulina hewitsonii (Butler, 1867)
SPEC Lazulina tolumnia (Cramer, 1777)

GENU Hermeuptychia Forster, 1964

SPEC Hermeuptychia atalanta (Butler, 1867)
SPEC Hermeuptychia camerta (Cramer, 1780)
SPEC Hermeuptychia canthe (Hübner, [1811])
SPEC Hermeuptychia clara Nakahara, Tan, Lamas & Willmott, 2016
SPEC Hermeuptychia cucullina (Weymer, 1911)
SPEC Hermeuptychia fallax (C. Felder & R. Felder, 1862)

SUBS Hermeuptychia fallax fallax (C. Felder & R. Felder, 1862)
SUBS Hermeuptychia fallax marinha Anken, 1994

SPEC Hermeuptychia gisella (Hayward, 1957)
SPEC Hermeuptychia harmonia (Butler, 1867)
SPEC Hermeuptychia hermes (Fabricius, 1775)
SPEC Hermeuptychia hermybius Grishin, 2014
SPEC Hermeuptychia intricata Grishin, 2014
SPEC Hermeuptychia lupita (Reakirt, [1867])
SPEC Hermeuptychia maimoune (Butler, 1870)
SPEC Hermeuptychia occidentalis Grishin, 2021
SPEC Hermeuptychia pimpla (C. Felder & R. Felder, 1862)
SPEC Hermeuptychia sinuosa Grishin, 2021
SPEC Hermeuptychia sosybius (Fabricius, 1793)
SPEC Hermeuptychia undulata (Butler, 1867)

GENU Saurona Huertas & Willmott, 2023

SPEC Saurona aurigera (Weymer, [1911])
SPEC Saurona triangula (Aurivillius, 1929)

GENU Pindis R. Felder, 1869

SPEC Pindis squamistriga R. Felder, 1869

GENU Amphidecta Butler, 1867

GSYN Rareuptychia Forster, 1964

SPEC Amphidecta calliomma (C. Felder & R. Felder, 1862)
SPEC Amphidecta clio (Weymer, 1911)
SPEC Amphidecta pignerator Butler, 1867

SUBS Amphidecta pignerator pignerator Butler, 1867
SUBS Amphidecta pignerator simplicia Weymer, 1910

SPEC Amphidecta reynoldsi Sharpe, 1890

Incertae sedis

SPEC Incertae_sedis insignis (Butler, 1867)
SPEC Incertae_sedis nerita (Capronnier, 1881)
SPEC Incertae_sedis ordinata (Weymer, 1911)
SPEC "Pharneuptychia" innocentia (C. Felder & R. Felder, 1867)

GENU Argentaria Huertas & Willmott, 2023

SPEC Argentaria argyropsacas (Bryk, 1953)
SPEC Argentaria clementia (Butler, 1877)
SPEC Argentaria clorimena (Stoll, 1790)

SSYN Splendeuptychia boliviensis Forster, 1964

SPEC Argentaria cosmophila (Hübner, 1823)
SPEC Argentaria hygina (Butler, 1877)
SPEC Argentaria itonis (Hewitson, 1862)
SPEC Argentaria jadea Brévignon & M. Benmesbah, 2012
SPEC Argentaria kendalli L. Miller, 1976
SPEC Argentaria libitina (Butler, 1870)
SPEC Argentaria pagyris (Godart, [1824])
SPEC Argentaria quadrina (Butler, 1869)
SPEC Argentaria salvini (Butler, 1867)
SPEC Argentaria telesphora (Butler, 1877)
SPEC Argentaria zischkai Forster, 1964

GENU Zischkaia Forster, 1964

SPEC Zischkaia abanico Nakahara & Petit, 2019
SPEC Zischkaia amalda (Weymer, 1911)
SPEC Zischkaia arctoa Nakahara, 2019
SPEC Zischkaia arenisca Nakahara, Willmott & Hall, 2019
SPEC Zischkaia argyrosflecha Nakahara, L. Miller & Huertas, 2019
SPEC Zischkaia baku Zacca, Dolibaina & Dias, 2019
SPEC Zischkaia chullachaki Nakahara & Zacca, 2019
SPEC Zischkaia josti Nakahara & Kleckner, 2019
SPEC Zischkaia mielkeorum Dolibaina, Dias & Zacca, 2019
SPEC Zischkaia pacarus (Godart, [1824])

SSYN Euptychia fumata Butler, 1867

SPEC Zischkaia saundersii (Butler, 1867)
SPEC Zischkaia warreni Dias, Zacca & Dolibaina, 2019

GENU Godartiana Forster, 1964

GSYN Cercyeuptychia Miller & Emmel 1971
GSYN Praefaunula Forster 1964

SPEC Godartiana amadoi M. Paluch, T. Zacca & A.V.L. Freitas, 2016
SPEC Godartiana armilla (Butler, 1867)
SPEC Godartiana astronesthes Lamas & Nakahara, 2018
SPEC Godartiana byses (Godart, [1824])
SPEC Godartiana luederwaldti (Spitz, 1931)
SPEC Godartiana muscosa (Butler, 1870)

GENU Forsterinaria Gray, 1973

GSYN Haywardina Forster 1964
GSYN Forsterina Lamas 1977
GSYN Forsterinonia Brown 1992
GSYN Guaianaza Freitas & Peña 2006

SPEC Forsterinaria anophthalma (C. Felder & R. Felder, 1867)
SPEC Forsterinaria antje Peña & Lamas, 2005
SPEC Forsterinaria boliviana (Godman, 1905)
SPEC Forsterinaria chaniorum Viloria & Costa, 2019
SPEC Forsterinaria coipa Peña & Lamas, 2005
SPEC Forsterinaria difficilis (Forster, 1964)
SPEC Forsterinaria emo Zubek, Pyrcz & Boyer, 2014
SPEC Forsterinaria enjuerma Peña & Lamas, 2005
SPEC Forsterinaria falcata Peña & Lamas, 2005
SPEC Forsterinaria guaniloi Peña & Lamas, 2005
SPEC Forsterinaria hannieri Zubek, Pyrcz & Boyer, 2011
SPEC Forsterinaria inornata (C. Felder & R. Felder, 1867)

SUBS Forsterinaria inornata inornata (C. Felder & R. Felder, 1867)
SUBS Forsterinaria inornata magdalena (Hayward, 1957)

SPEC Forsterinaria itatiaia Peña & Lamas, 2005
SPEC Forsterinaria necys (Godart, [1824])
SPEC Forsterinaria neonympha (C. Felder & R. Felder, 1867)
SPEC Forsterinaria pallida Peña & Lamas, 2005

SUBS Forsterinaria pallida pallida Peña & Lamas, 2005
SUBS Forsterinaria pallida aurita Peña & Lamas, 2005

SPEC Forsterinaria pichita Peña & Lamas, 2005
SPEC Forsterinaria pilosa Peña & Lamas, 2005
SPEC Forsterinaria pronophila (Butler, 1867)
SPEC Forsterinaria proxima (Hayward, 1957)
SPEC Forsterinaria pseudinornata (Forster, 1964)
SPEC Forsterinaria punctata Peña & Lamas, 2005
SPEC Forsterinaria pyrczi Peña & Lamas, 2005
SPEC Forsterinaria quantius (Godart, [1824])
SPEC Forsterinaria rotunda Peña & Lamas, 2005
SPEC Forsterinaria rustica (Butler, 1868)

SUBS Forsterinaria rustica rustica (Butler, 1868)
SUBS Forsterinaria rustica villarresi (Dognin, 1887)
SUBS Forsterinaria rustica glendita Peña & Lamas, 2005

SPEC Forsterinaria stella (Hayward, 1957)

GENU Harjesia Forster, 1964

SPEC Harjesia argentata Nakahara, Zacca and Lamas, 2018
SPEC Harjesia blanda (Möschler, 1877)
SPEC Harjesia obscura (Butler, 1867)

GENU Orotaygetis Nakahara & Zacca, 2018

SPEC Orotaygetis surui Nakahara, Zacca & Lamas, 2018

GENU Parataygetis Forster, 1964

SPEC Parataygetis albinotata (Butler, 1867)
SPEC Parataygetis lineata (Godman & Salvin, 1880)

GENU Posttaygetis Forster, 1964

SPEC Posttaygetis penelea (Cramer, 1777)

GENU Pseudodebis Forster, 1964

GSYN Taygetomorpha L. D. Miller, 2004

SPEC Pseudodebis celia (Cramer, 1779)
SPEC Pseudodebis darrenthroopi Nakahara & Willmott, 2022
SPEC Pseudodebis dubiosa Forster, 1964
SPEC Pseudodebis euptychidia (Butler, 1868)
SPEC Pseudodebis hartmanni Nakahara & MacDonald, 2021
SPEC Pseudodebis marpessa (Hewitson, 1862)
SPEC Pseudodebis nakamurai Nakahara & Willmott, 2021
SPEC Pseudodebis pieti Nakahara & Willmott, 2021
SPEC Pseudodebis puritana (Weeks, 1902)
SPEC Pseudodebis tigrillo Nakahara & Willmott, 2022
SPEC Pseudodebis valentina (Cramer, 1779)
SPEC Pseudodebis vrazi (Kheil, 1896)
SPEC Pseudodebis zimri (Butler, 1869)

GENU Taguaiba Freitas, Zacca & Siewert, 2023

SPEC Taguaiba drogoni (Siewert, Zacca, Dias & Freitas, 2013)
SPEC Taguaiba fulginia (R.F. d'Almeida, 1922)
SPEC Taguaiba rectifascia (Weymer, 1907)
SPEC Taguaiba servius (Weymer, 1910)
SPEC Taguaiba ypthima (Hübner, [1821])

GENU Sepona Freitas & Barbosa, 2015

SPEC Sepona punctata (Weymer, 1911)

GENU Taygetina Forster, 1964

GSYN Coeruleotaygetis Forster, 1964

SPEC Taygetina accacioi Nakahara & Freitas, 2019
SPEC Taygetina banghaasi (Weymer, 1910)
SPEC Taygetina brocki Lamas & Nakahara, 2019
SPEC Taygetina gulnare (Butler, 1870)
SPEC Taygetina kerea (Butler, 1869)
SPEC Taygetina oreba (Butler, 1870)
SPEC Taygetina peribaea (Godman & Salvin, 1880)
SPEC Taygetina weymeri (Draudt, 1912)

GENU Taygetis Hübner, [1819]

SPEC Taygetis acuta Weymer, 1910
SPEC Taygetis angulosa Weymer, 1907
SPEC Taygetis asterie Weymer, 1910
SPEC Taygetis chiquitana Forster, 1964
SPEC Taygetis chrysogone E. Doubleday, [1849]
SPEC Taygetis cleopatra C. Felder & R. Felder, 1867
SPEC Taygetis echo (Cramer, 1775)

SUBS Taygetis echo echo (Cramer, 1775)
SUBS Taygetis echo koepckei Forster, 1964
SUBS Taygetis echo velutina Staudinger, [1887]

SPEC Taygetis elegia Weymer, 1910
SPEC Taygetis inambari L. Miller & Lamas, 1999
SPEC Taygetis inconspicua Draudt, 1931
SPEC Taygetis kharisma Brévignon & M. Benmesbah, 2011
SPEC Taygetis laches (Fabricius, 1793)

SUBS Taygetis laches laches (Fabricius, 1793)
SUBS Taygetis laches marginata Staudinger, [1887]

SPEC Taygetis larua C. Felder & R. Felder, 1867

SSYN Taygetis armillata Butler, 1868

SPEC Taygetis leuctra Butler, 1870
SPEC Taygetis mermeria (Cramer, 1776)

SUBS Taygetis mermeria mermeria (Cramer, 1776)
SUBS Taygetis mermeria excavata Butler, 1868
SUBS Taygetis mermeria griseomarginata L. Miller, 1978

SPEC Taygetis oyapock Brévignon, 2007
SPEC Taygetis rufomarginata Staudinger, 1888

SUBS Taygetis rufomarginata rufomarginata Staudinger, 1888
SUBS Taygetis rufomarginata tapulunmi Brévignon, 2005

SPEC Taygetis sosis Hopffer, 1874

SSYN Taygetis godmani Weymer, 1910

SPEC Taygetis sylvia H. Bates, 1866
SPEC Taygetis thamyra (Cramer, 1779)

SSYN Taygetis magna D’Almeida, 1922
SSYN Taygetis isis Bargmann, 1928
SSYN Taygetis stollei Ribeiro, 1931

SPEC Taygetis tripunctata Weymer, 1907
SPEC Taygetis uncinata Weymer, 1907
SPEC Taygetis uzza Butler, 1869

SSYN Taygetis salvini Staudinger, [1887]

SPEC Taygetis virgilia (Cramer, 1776)

SSYN Papilio rebecca Fabricius, 1793
SSYN Taygetis nympha Butler, 1868
SSYN Taygetis erubescens Butler, 1868
SSYN Taygetis daguana Bargmann, 1928
SSYN Taygetis pseudorufomarginata Bargmann, 1929

SPEC Taygetis zippora Butler, 1869

GENU Chloreuptychia Forster, 1964

SPEC Chloreuptychia agatha (Butler, 1867)
SPEC Chloreuptychia amethysta Brévignon & M. Benmesbah, 2011
SPEC Chloreuptychia chlorimene (Hübner, [1819])
SPEC Chloreuptychia gordonsmalli Thurman, MacDonald & Nakahara, 2021

GENU Archeuptychia Forster, 1964

SPEC Archeuptychia cluena (Drury, 1782)

GENU Stephenympha Viloria, 2022

SPEC Stephenympha pauliana Viloria, 2022
SPEC Stephenympha arius (Weymer, 1911)
SPEC Stephenympha eriphule (Butler, 1867)

GENU Cisandina Nakahara & Espeland, 2022

SPEC Cisandina castanya Lamas & Nakahara, 2022
SPEC Cisandina esmeralda Nakahara & Barbosa, 2022
SPEC Cisandina fida (Weymer, 1911)

SUBS Cisandina fida fida (Weymer, 1911)
SUBS Cisandina fida directa Nakahara & Willmott, 2022
SPEC Cisandina lea (Cramer, 1777)
SPEC Cisandina philippa (Butler, 1867)
SPEC Cisandina sanmarcos (Nakahara & Lamas, 2018)
SPEC Cisandina trinitensis (Brévignon & Benmesbah, 2011)

GENU Pseudeuptychia Forster, 1964

SPEC Pseudeuptychia callichloris (Butler, 1867)
SPEC Pseudeuptychia cuzquenya Nakahara & Lamas, 2018
SPEC Pseudeuptychia hemileuca (Staudinger, [1886])
SPEC Pseudeuptychia herseis (Godart, [1824])
SPEC Pseudeuptychia languida (Butler, 1871)

SUBS Pseudeuptychia languida languida (Butler, 1871)
SUBS Pseudeuptychia languida austrina Nakahara & Lamas, 2018

SPEC Pseudeuptychia marica (Weymer, 1911)
SPEC Pseudeuptychia rectilinea (Brévignon, Rosant, Lamas & Willmott, 2019)

GENU Amiga Nakahara, Willmott & Espeland, 2019

SPEC Amiga arnaca (Fabricius, 1776)

SUBS Amiga arnaca arnaca (Fabricius, 1776)
SUBS Amiga arnaca adela Nakahara & Espeland, 2019
SUBS Amiga arnaca indianacristoi Nakahara & Marín, 2019

SPEC Amiga sericeella (Bates, 1864)

GENU Erichthodes Forster, 1964

SPEC Erichthodes antonina (C. Felder & R. Felder, 1867)

Removed from Erichthodes, unplaced for the moment (Espeland et al., 2023)

SPEC "Erichthodes" eremita Lamas, Willmott & Radford, 2018
SPEC "Erichthodes" jovita (C. Felder & R. Felder, 1867)
SPEC "Erichthodes" julia (Weymer, 1911)

GENU Euptychoides Forster, 1964

SPEC Euptychoides albofasciata (Hewitson, 1869)
SPEC Euptychoides inani (Staudinger, [1886])
SPEC Euptychoides laccine (C. Felder & R. Felder, 1867)
SPEC Euptychoides pseudosaturnus Forster, 1964

GENU Xenovena Marín & Nakahara, 2023

SPEC Xenovena murrayae (Brévignon, 2005)

GENU Megeuptychia Forster, 1964

SPEC Megeuptychia antonoe (Cramer, 1775)
SPEC Megeuptychia monopunctata Willmott & Hall, 1995
SPEC Megeuptychia souadae (M. Benmesbah, 2015)

GENU Neonympha Hübner, 1818

SPEC Neonympha areolatus (J. E. Smith, 1797)

SUBS Neonympha areolatus areolatus (J. E. Smith, 1797)

SSYN Papilio phocion Fabricius, 1793
SSYN Neonympha helicta (Hübner, [1808])

SUBS Neonympha areolatus septentrionalis W. Davis, 1924
SUBS Neonympha areolatus dadeensis Gatrelle, 1999

SPEC Neonympha mitchellii French, 1889

SUBS Neonympha mitchellii mitchellii French, 1889
SUBS Neonympha mitchellii francisci Parshall & Kral, 1989

GENU Nhambikuara Freitas, Barbosa & Zacca, 2018

SPEC Nhambikuara ackeryi (Huertas, Rios & LeCrom, 2009)
SPEC Nhambikuara cerradensis Freitas, Barbosa & Zacca, 2018
SPEC Nhambikuara doxes (Godart, [1824])
SPEC Nhambikuara erycina (Butler, 1867)
SPEC Nhambikuara furina (Hewitson, 1862)
SPEC Nhambikuara gemmula (Butler, 1867)
SPEC Nhambikuara junonia (Butler, 1867)
SPEC Nhambikuara latia (Butler, 1867)
SPEC Nhambikuara mercedes (Huertas, 2011)
SPEC Nhambikuara mima (Butler, 1867)
SPEC Nhambikuara toynei (Willmott & J. Hall, 1995)

GENU Optimandes Marín, Nakahara & Willmott, 2019

SPEC Optimandes eugenia (C. Felder & R. Felder, 1867)

SUBS Optimandes eugenia eugenia (C. Felder & R. Felder, 1867)

SSYN Euptychia phineus Butler, 1867

SUBS Optimandes eugenia transversa (Weymer, 1911)

SPEC Optimandes mocha Willmott, Hall & Lamas, 2019

GENU Pareuptychia Forster, 1964

SPEC Pareuptychia binocula (Butler, 1869)
SPEC Pareuptychia difficilis Forster, 1964
SPEC Pareuptychia hervei Brévignon, 2005
SPEC Pareuptychia hesionides Forster, 1964

SUBS Pareuptychia hesionides hesionides Forster, 1964
SUBS Pareuptychia hesionides deviae Brévignon, 2005

SPEC Pareuptychia lydia (Cramer, 1777)
SPEC Pareuptychia metaleuca (Boisduval, 1870)
SPEC Pareuptychia milleri Nakahara, Marín & Neild, 2016
SPEC Pareuptychia ocirrhoe (Fabricius, 1776)

SUBS Pareuptychia ocirrhoe ocirrhoe (Fabricius, 1776)
SUBS Pareuptychia ocirrhoe interjecta (R.F. d'Almeida, 1952)

SPEC Pareuptychia summandosa (Gosse, 1880)

GENU Satyrotaygetis Forster, 1964

GSYN Macrocissia Viloria, Le Crom & Andrade, 2019

SPEC Satyrotaygetis iris (C. Felder & R. Felder, 1867)

SSYN Euptychia ayaya Butler, 1867

SPEC Satyrotaygetis satyrina (H. Bates, 1865)

SSYN Satyrotaygetis gigas (Butler, 1867)
SSYN Satyrotaygetis incerta (Butler & Druce, 1872)

SPEC Satyrotaygetis tiessa (Hewitson, 1869)

GENU Taydebis Freitas, 2003

GSYN Prenda Freitas & Mielke, 2011

SPEC Taydebis melobosis (Capronnier, 1874)

SSYN Taydebis peculiaris (Butler, 1874)

SPEC Taydebis clarissa (Freitas & Mielke, 2011)
SPEC Taydebis guria Zacca, Casagrande & Mielke, 2021

GENU Caeruleuptychia Forster, 1964

GSYN Weymerana Forster 1964

SPEC Caeruleuptychia aegrota (Butler, 1867)
SPEC Caeruleuptychia aetherialis (Butler, 1877)
SPEC Caeruleuptychia coelestis (Butler, 1867)
SPEC Caeruleuptychia coelica (Hewitson, 1869)
SPEC Caeruleuptychia cyanites (Butler, 1871)

SSYN Euptychia stigmatica Godman, 1905

SPEC Caeruleuptychia harrisi Nakahara & Freitas, 2023
SPEC Caeruleuptychia aemulatio Nakahara & Willmott, 2023
SPEC Caeruleuptychia divina (Weymer, 1911)
SPEC Caeruleuptychia francisca (Butler, 1870)
SPEC Caeruleuptychia glauca (Weymer, 1911)
SPEC Caeruleuptychia helena (Anken, 1994)
SPEC Caeruleuptychia helios (Weymer, 1911)

SSYN Caeruleuptychia helios caelestissima Brévignon, 2010
SSYN Magneuptychia keltoumae Brévignon & Benmesbah, 2012

SPEC Caeruleuptychia lobelia (Butler, 1870)
SPEC Caeruleuptychia mare (Butler, 1869)
SPEC Caeruleuptychia maryzenderae Lamas & Nakahara, 2017
SPEC Caeruleuptychia penicillata (Godman, 1905)
SPEC Caeruleuptychia pilata (Butler, 1867)
SPEC Caeruleuptychia romani (Aurivillius, 1929)
SPEC Caeruleuptychia saul Brévignon & M. Benmesbah, 2011
SPEC Caeruleuptychia scopulata (Godman, 1905)
SPEC Caeruleuptychia scripta Nakahara, T. Zacca & B. Huertas, 2017
SPEC Caeruleuptychia tenera (Weymer, 1911)
SPEC Caeruleuptychia thaliana Nakahara & Piovesan, 2022
SPEC Caeruleuptychia trembathi Willmott, Nakahara, J. Hall & Neild, 2017
SPEC Caeruleuptychia twalela Brévignon, 2005
SPEC Caeruleuptychia umbrosa (Butler, 1870)
SPEC Caeruleuptychia urania (Butler, 1867)

SSYN Euptychia caerulea Butler, 1869
SSYN Caeruleuptychia sinchi Henao-Bañol, 2019

GENU Capronnieria Forster, 1964

SPEC Capronnieria galesus (Godart, [1824])
SPEC Capronnieria narapa (Schaus, 1902)

GENU Cepheuptychia Forster, 1964

SPEC Cepheuptychia angelica (Butler, 1874)
SPEC Cepheuptychia cephus (Fabricius, 1775)
SPEC Cepheuptychia glaucina (Bates, 1864)

GENU Colombeia Viloria, Andrade & Le Crom, 2019

SPEC Colombeia hotchkissi (Dyar, 1913)
SPEC Colombeia mycalesis (Röber, 1927)
SPEC Colombeia nossis (Hewitson, 1862)

GENU Deltaya Willmott, Nakahara & Espeland, 2023

SPEC Deltaya andrei (Zacca, Casagrande & Mielke, 2017)
SPEC Deltaya louisammour (Benmesbah & Zacca, 2018)
SPEC Deltaya ocypete (Fabricius, 1776)
SPEC Deltaya opima (Weymer, 1911)
SPEC Deltaya pallema (Schaus, 1902)
SPEC Deltaya probata (Weymer, 1911)

GENU Emeryus Zacca, Casagrande & Mielke, 2020

SPEC Emeryus argulus (Godart, [1824])

SUBS Emeryus argulus argulus (Godart, [1824])
SUBS Emeryus argulus magnum Zacca, Casagrande & Mielke, 2020

SPEC Emeryus difficilis (Forster, 1964)
SPEC Emeryus numeria (C. Felder & R. Felder, 1867)

GENU Huberonympha Viloria & Costa, 2016

SPEC Huberonympha neildi Viloria, Costa, Fratello & Nakahara, 2016

GENU Magneuptychia Forster, 1964

SPEC Magneuptychia lethra (Möschler, 1883)
SPEC Magneuptychia libye (Linnaeus, 1767)

GENU Modica Zacca, Casagrande & Willmott, 2023

SPEC Modica confusa (Staudinger, 1887)
SPEC Modica fugitiva (Lamas, [1997])
SPEC Modica kamel (Benmesbah & Zacca, 2018)
SPEC Modica maripa (Brévignon, 2005)
SPEC Modica myncea (Cramer, 1780)

GENU Occulta Nakahara & Willmott, 2023

SPEC Occulta ocnus (Butler, 1867)

GENU Trico Nakahara & Espeland, 2023

SPEC Trico tricolor (Hewitson, 1850)

SSUB Trico tricolor tricolor (Hewitson, 1850)
SSUB Trico tricolor fulgora (Butler, 1869)

GENU Malaveria Viloria & Benmesbah, 2021

GSYN Rudyphthimoides Viloria, 2022

SPEC Malaveria affinis (Butler, 1867)
SPEC Malaveria alcinoe (C. Felder & R. Felder, 1867)
SPEC Malaveria argyrospila (Butler, 1867)
SPEC Malaveria ballofi Benmesbah & Viloria, 2021
SPEC Malaveria bottoi Benmesbah & Viloria, 2021
SPEC Malaveria duponti Benmesbah & Murienne, 2021
SPEC Malaveria erigone (Butler, 1867)
SPEC Malaveria grimon (Godart, [1824])
SPEC Malaveria guenzeli (Anken, 1994)
SPEC Malaveria iseai (Viloria, 2022)
SPEC Malaveria maepius (Godart, [1824])

SSYN Euptychia perfuscata Butler, 1867

SPEC Malaveria mimas (Godman, 1905)
SPEC Malaveria mimula (Hayward, 1954)
SPEC Malaveria mythra (Weymer, 1911)
SPEC Malaveria nebulosa (Butler, 1867)
SPEC Malaveria neomaenas (Hayward, 1967)
SPEC Malaveria rodriguezi Benmesbah & Viloria, 2020

SUBS Malaveria rodriguezi rodriguezi Benmesbah & Viloria, 2020
SUBS Malaveria rodriguezi risaralda Benmesbah & Viloria, 2020

GENU Modestia Viloria & Benmesbah, 2020

SPEC Modestia agnata (Schaus, 1913)
SPEC Modestia analis (Godman, 1905)
SPEC Modestia drymo (Schaus, 1913)
SPEC Modestia gomezi (Singer, DeVries & P. Ehrlich, 1983)
SPEC Modestia harpyia (C. Felder & R. Felder, 1867)

SUBS Modestia harpyia harpyia (C. Felder & R. Felder, 1867)
SUBS Modestia harpyia batesii (Butler, 1867)

SPEC Modestia modesta (Butler, 1867)
SPEC Modestia remypignoux Benmesbah & Viloria, 2020

SUBS Modestia remypignoux remypignoux Benmesbah & Viloria, 2020
SUBS Modestia remypignoux shueyi Benmesbah & Viloria, 2020

SPEC Modestia sylvina (C. Felder & R. Felder, 1867)

GENU Omacha Andrade, Viloria, Henao & Le Crom, 2019

SPEC Omacha pax (Huertas, Lamas, Fagua & Willmott, 2016)

GENU Paryphthimoides Forster, 1964

SPEC Paryphthimoides brixiola (Butler, 1867)
SPEC Paryphthimoides brixius (Godart, [1824])

SUBS Paryphthimoides brixius brixius (Godart, [1824])
SUBS Paryphthimoides brixius madre Zacca, Mielke, Casagrande & Lamas, 2020
SUBS Paryphthimoides brixius rouranensis Brévignon, 2020

SPEC Paryphthimoides flavofascia (Zacca & Siewert, 2014)
SPEC Paryphthimoides fridae Zacca, Casagrande, Huertas & Nakahara, 2020
SPEC Paryphthimoides jorupe Zacca, Casagrande, Checa & Willmott, 2020
SPEC Paryphthimoides joyceae Singer, DeVries & P. Ehrlich, 1983
SPEC Paryphthimoides poltys (Prittwitz, 1865)

SUBS Paryphthimoides poltys poltys (Prittwitz, 1865)

SSYN Paryphthimoides zeredatha (Butler, 1869)
SSYN Paryphthimoides bahneri Anken, 1994
SSYN Paryphthimoides nerita (Capronnier, 1881)

SUBS Paryphthimoides poltys numilia (C. Felder & R. Felder, 1867)

SSYN Euptychia binalinea Butler, 1867
SSYN Paryphthimoides poltys bahiana Bryk, 1953

SPEC Paryphthimoides pseudoconfusa Singer, DeVries & P. Ehrlich, 1983
SPEC Paryphthimoides sheba Brévignon & M. Benmesbah, 2012
SPEC Paryphthimoides terrestris (Butler, 1867)

SUBS Paryphthimoides terrestris terrestris (Butler, 1867)
SUBS Paryphthimoides terrestris muyrakytan Zacca, Casagrande & Mielke, 2020
SUBS Paryphthimoides terrestris araguaianus Zacca, Casagrande & Mielke, 2020
SUBS Paryphthimoides terrestris grevei Zacca, Casagrande & Mielke, 2020

SPEC Paryphthimoides touloulou (Benmesbah, 2015)
SPEC Paryphthimoides vestigiata (Butler, 1867)

GENU Scriptor Nakahara & Espeland, 2020

SPEC Scriptor sphenophorus Lamas & Nakahara, 2020

GENU Splendeuptychia Forster, 1964

GSYN Nubila Viloria, Andrade & Henao, 2019

SPEC Splendeuptychia ambra (Weymer, [1911])
SPEC Splendeuptychia ashna (Hewitson, 1869)
SPEC Splendeuptychia gera (Hewitson, 1850)
SPEC Splendeuptychia metagera (Butler, 1867)
SPEC Splendeuptychia moderata (Weymer, 1911)
SPEC Splendeuptychia nortia (Hewitson, 1862)

SSYN Magneuptychia nobilis (Weymer, 1911)

SPEC Splendeuptychia purusana (Aurivillius, 1929)
SPEC Splendeuptychia segesta (Weymer, 1911)
SPEC Splendeuptychia tupinamba Freitas, Huertas & Rosa, 2021

GENU Stevenaria Viloria, Costa, Neild & Nakahara, 2016

SPEC Stevenaria divergens (Butler, 1867)
SPEC Stevenaria nakaharai Viloria, Costa, S. Fratello & Neild, 2016
SPEC Stevenaria yutajensis Viloria & Costa, 2016

GENU Vareuptychia Forster 1964

SPEC Vareuptychia similis (Butler, 1867)
SPEC Vareuptychia themis (Butler, 1867)

GENU Llorenteana Viloria & Martinez, 2019

SPEC Llorenteana pellonia (Godman, 1901)

---

Sources:

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