Satyrina

Niklas Wahlberg

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Updated 2022-09-24. Based on

• Markku Savela's pages
• The Butterflies of America
• Kleckova, I., Cesanek, M., Fric, Z. & Pellissier, L. 2015: Diversification of the cold-adapted butterfly genus Oeneis related to Holarctic biogeography and climatic niche shifts. Molecular Phylogenetics and Evolution 92: 255-265. doi:10.1016/j.ympev.2015.06.012
• Lukhtanov, V. A. & Dubatolov, V. V. 2020: Phylogenetic position and taxonomic rearrangement of Davidina (Lepidoptera: Nymphalidae), an enigmatic butterfly genus new for Europe and America. Zoological Journal of the Linnean Society 190: 1036–1053. doi:10.1093/zoolinnean/zlaa104
• Russell PJC, Vane-Wright RI 2022: Papilio hermione Linnaeus, type species of Hipparchia Fabricius (Lepidoptera, Satyrinae): restoring stability to the application of these names. Nota Lepidopterologica 45: 279-294. doi:10.3897/nl.45.85341

SUBT Satyrina

GENU Argestina Riley, 1923

SPEC Argestina phantasta Goltz, 1938
SPEC Argestina pomena (Evans, 1915)
SPEC Argestina inconstans (South, 1913)
SPEC Argestina irma (Evans, 1923)
SPEC Argestina karta (Riley, 1923)
SPEC Argestina nitida (Riley, 1923)
SPEC Argestina waltoni (Elwes, 1906)

ORIG Callerebia waltoni Elwes, 1906

SUBS Argestina waltoni waltoni (Elwes, 1906)
SUBS Argestina waltoni pseudonitida Huang, 1998

GENU Berberia de Lesse, 1951

SPEC Berberia abdelkader (Pierret, 1837)

ORIG Satyrus Abd-el-Kader Pierret, 1837

SUBS Berberia abdelkader abdelkader (Pierret, 1837)

SSYN Berberia abdelkader alfae Slaby, 1981
SSYN Berberia abdelkader saharae Slaby, 1981
SSYN Berberia abdelkader arvorum Slaby, 1981

SUBS Berberia abdelkader nelvai (Seitz, 1911)

ORIG Satyrus abdelkader nelvai Seitz, 1911
SSYN Berberia abdelkader marteni Chnéour, 1935

SUBS Berberia abdelkader taghzefti (Wyatt, 1952)

ORIG Satyrus abdelkader taghzefti Wyatt, 1952

SPEC Berberia lambessanus (Staudinger, 1901)

ORIG Satyrus abdelkader var. lambessanus Staudinger, 1901
SSYN Satyrus (Cercyonis) abdelkader romeii Rothschild, 1933
SSYN Satyrus abdelkader alexander Chnéour, 1937

GENU Hipparchia Fabricius, 1807

GSYN Eumenis Hübner, 1819
GSYN Nytha Billberg, 1820
GSYN Melania Sodoffsky, 1837 (unn. repl. Hipparchia Fabricius, 1807, preocc. Melania Lamarck, 1799, Melania Perry, 1811)
GSYN Pseudotergumia Agenjo, 1947
GSYN Neohipparchia de Lesse, 1951

SPEC Hipparchia fagi (Scopoli, 1763)

ORIG Papilio fagi Scopoli, 1763
SSYN Papilio hermione Linnaeus, 1764

SUBS Hipparchia fagi fagi (Scopoli, 1763)
SUBS Hipparchia fagi tetrica Fruhstorfer, 1907

SPEC Hipparchia alcyone (Denis & Schiffermüller, 1775)
SPEC Hipparchia syriaca (Staudinger, 1871)
SPEC Hipparchia autonoe (Esper, 1784)

ORIG Papilio autonoe Esper, 1784

SUBS Hipparchia autonoe autonoe (Esper, 1784)
SUBS Hipparchia autonoe orchomenus (Fruhstorfer, 1911)
SUBS Hipparchia autonoe sibirica (Staudinger, 1861)
SUBS Hipparchia autonoe maxima Bang-Haas
SUBS Hipparchia autonoe wutaiensis Murayama

SPEC Hipparchia neomiris (Godart, 1824)
SPEC Hipparchia aristaeus (Bonelli, 1826)

ORIG Papilio Satyrus aristaeus Bonelli, 1826
SSYN Hipparchia aristaues blachieri (Fruhstorfer, 1908)
SSYN Hipparchia aristause ballettoi (Kudrna, 1984)

SUBS Hipparchia aristaeus aristaeus (Bonelli, 1826)
SUBS Hipparchia aristaeus algirica (Oberthür, 1876)

SPEC Hipparchia senthes (Fruhstorfer, 1908)
SPEC Hipparchia sborbonii Kudrna, 1984
SPEC Hipparchia maderensis (Bethune-Baker, 1891)
SPEC Hipparchia cretica (Rebel, 1916)
SPEC Hipparchia semele (Linnaeus, 1758)

ORIG Papilio semele (Linnaeus, 1758)
SSYN Hipparchia semele leighebi Kudrna, 1976
SSYN Hipparchia semele cadmu Fruhstorfer, 1908

SPEC Hipparchia mersina (Staudinger, 1871)

SSYN Hipparchia malickyi Kudrna, 1976

SPEC Hipparchia christenseni Kudrna, 1976
SPEC Hipparchia volgensis (Mazochin-Porshnjakov, 1952)
SPEC Hipparchia pellucida (Stauder, 1924)

SUBS Hipparchia pellucida pellucida (Stauder, 1924)
SUBS Hipparchia pellucida turkmenica (Heydemann, 1942)

SPEC Hipparchia statilinus (Hufnagel, 1766)

ORIG Papilio statilinus Hufnagel, 1766

SUBS Hipparchia statilinus statilinus (Hufnagel, 1766)
SUBS Hipparchia statilinus sylvicola (Austaut, 1880)

ORIG Satyrus sylvicola Austaut, 1880
SSYN Satyrus fauna var. cinerea Oberthür, 1907
SSYN Eumenis statilinus anaxarchus Fruhstorfer, 1909
SSYN Satyrus sylvicola oberthueri Rothschild, 1917
SSYN Satyrus sylvicola cordieri Chnéour, 1950
SSYN Hipparchia sylvicola rungsi Varin, 1952

SPEC Hipparchia fatua (Feryer, 1844)

SUBS Hipparchia fatua fatua (Feryer, 1844)
SUBS Hipparchia fatua persiscana (Verity, 1937)
SUBS Hipparchia fatua klapperichi (Gross & Ebert, 1975)

SPEC Hipparchia parisatis (Kollar, 1849)

ORIG Eumenis parisatis Kollar, 1849
SSYN Satyrus parisatis (Kollar, 1849)

SUBS Hipparchia parisatis parisatis (Kollar, 1849)
SUBS Hipparchia parisatis xizangensis (Chou, 1994)
SUBS Hipparchia parisatis laeta (Christoph, 1877)
SUBS Hipparchia parisatis macrophthalma Eversmann, 1851

SPEC Hipparchia stulta (Staudinger, 1882)
SPEC Hipparchia ellena (Oberthür, 1894)

SUBS Hipparchia ellena ellena (Oberthür, 1894)
SUBS Hipparchia ellena caroli (Rothschild, 1933)

SPEC Hipparchia wyssii (Christ, 1889)
SPEC Hipparchia bacchus Higgins, 1967
SPEC Hipparchia gomera Higgins, 1967
SPEC Hipparchia azorina (Strecker, 1899)
SPEC Hipparchia hansii (Austaut, 1879)

ORIG Satyrus Hansii Austaut, 1879
SSYN Satyrus Fauna-holli Oberthür, 1909
SSYN Satyrus colombati Oberthür, 1921
SSYN Satyrus belouini Oberthür, 1921
SSYN Satyrus colombati ab. stellifer Chnéour, 1942
SSYN Satyrus statilinus f. chpakowskyi Chnéour, 1942
SSYN Satyrus hansii intermedia Chnéour, 1963
SSYN Satyrus hansii tlemceni Slaby, 1977
SSYN Hipparchia hansii tansleyi Tarrier, 1995
SSYN Neohipparchia hansii (Austaut, 1879)

SPEC Hipparchia powelli (Oberthür, 1910)

ORIG Satyrus powelli Oberthür, 1910

SPEC Hipparchia fidia (Linnaeus, 1767)

ORIG Papilio fidia Linnaeus, 1767
SSYN Satyrus fidia hebitis Rothschild, 1917
SSYN Satyrus fidia intermedia Rothschild, 1917
SSYN Satyrus fidia minerva Hemming, 1934
SSYN Hipparchia fidia kandarica Varin, 1954
SSYN Hipparchia fidia benimguildi Varin, 1954

SPEC Hipparchia pisidice Klug, 1832
SPEC Hipparchia tewfiki (Wiltshire, 1949)
SPEC Hipparchia delattini Kudrna, 1975
SPEC Hipparchia cypriensis (Holik, 1949)
SPEC Hipparchia tilosi (Manil, 1984)

GENU Kanetisa Moore, 1893

SPEC Kanetisa digna (Marshall, 1883)

ORIG Hipparchia digna Marshall, 1883

GENU Aulocera Butler, 1867

SPEC Aulocera sybillina Oberthür, 1890

SSYN Satyrus sybillina (Oberthür, 1890)

SPEC Aulocera brahminus (Blanchard, 1853)

ORIG Satyrus brahminus Blanchard, 1853

SPEC Aulocera brahminoides (Moore, 1896)
SPEC Aulocera magica (Oberthür, 1886)

SUBS Aulocera magica magica (Oberthür, 1886)
SUBS Aulocera magica amida Gross, 1958

SPEC Aulocera merlina (Oberthür, 1890)

SUBS Aulocera merlina merlina Oberthür, 1890
SUBS Aulocera merlina pulcheristriata Huang, 2001

SPEC Aulocera padma (Kollar, 1844)

ORIG Satyrus padma Kollar, 1844

SPEC Aulocera loha Doherty, 1886

SUBS Aulocera loha loha Doherty, 1886
SUBS Aulocera loha fulva Evans, 1923

SPEC Aulocera asahi Fujioka, 1970
SPEC Aulocera swaha (Kollar, 1844)

ORIG Satyrus swaha Kollar, 1844

SPEC Aulocera saraswati (Kollar, 1844)

ORIG Satyrus saraswati Kollar, 1844

SUBS Aulocera saraswati saraswati (Kollar, 1844)
SUBS Aulocera saraswati vishnu Gross, 1958
SUBS Aulocera saraswati chayuensis Huang, 2001

GENU Karanasa Moore, 1893

SPEC Karanasa voigti Bang-Haas, 1927

SUBS Karanasa voigti voigti Bang-Haas, 1927
SUBS Karanasa voigti nigrocellata Avinoff & Sweadner, 1951

SPEC Karanasa bolorica (Grum-Grshimailo, 1888)

ORIG Satyrus boloricus Grum-Grshimailo, 1888

SUBS Karanasa boloricus boloricus (Grum-Grshimailo, 1888)
SUBS Karanasa boloricus chitralica Tytler, 1926
SUBS Karanasa boloricus hodja Avinoff & Sweadner, 1951

SPEC Karanasa mohsenii Wyatt & Omoto, 1966
SPEC Karanasa decolorata (Staudinger, 1901)

SUBS Karanasa decolorata decolorata (Staudinger, 1901)
SUBS Karanasa decolorata grumi Avinoff & Sweadner, 1951
SUBS Karanasa decolorata mushketovi Avinoff & Sweadner, 1951

SSYN Karanasa decolorata roborovskyi Avinoff & Sweadner, 1951

SPEC Karanasa maureri Avinoff & Sweadner, 1951

SUBS Karanasa maureri maureri Avinoff & Sweadner, 1951
SUBS Karanasa maureri iskander Avinoff & Sweadner, 1951

SSYN Karanasa decolorata maidana Avinoff & Sweadner, 1951
SSYN Karanasa anzobica J.J. Shchetkin, 1986

SPEC Karanasa josephi (Staudinger, 1882)

ORIG Satyrus josephi Staudinger, 1882

SUBS Karanasa josephi josephi (Staudinger, 1882)
SUBS Karanasa josephi dissoluta (Staudinger, 1886)
SUBS Karanasa josephi darwasica Avinoff & Sweadner, 1951
SUBS Karanasa josephi oshanini Avinoff & Sweadner, 1951
SUBS Karanasa josephi jakobsoni Avinoff & Sweadner, 1951
SUBS Karanasa josephi hissariensis Avinoff & Sweadner, 1951

SPEC Karanasa talastauana O. Bang-Haas, 1927

ORIG Satyrus huebneri talastauana O. Bang-Haas, 1927

SUBS Karanasa talastauana talastauna O. Bang-Haas, 1927
SUBS Karanasa talastauana arasana Avinoff & Sweadner, 1951
SUBS Karanasa talastauana angrena Avinoff & Sweadner, 1951

SPEC Karanasa praestans Avinoff & Sweadner, 1951

SUBS Karanasa praestans praestans Avinoff & Sweadner, 1951
SUBS Karanasa praestans kasak Avinoff & Sweadner, 1951

SPEC Karanasa wilkinsi (Erschoff, 1884)

ORIG Satyrus wilkinsi Erschoff, 1884

SUBS Karanasa wilkinsi wilkinsi (Erschoff, 1884)
SUBS Karanasa wilkinsi robusta Avinoff & Sweadner, 1951
SUBS Karanasa wilkinsi dublitzkyi (O. Bang-Haas, 1927)
SUBS Karanasa wilkinsi durana Avinoff & Sweadner, 1951

SPEC Karanasa kirgizorum Avinoff & Sweadner, 1951
SPEC Karanasa leechi (Grum-Grshimailo, 1890)

ORIG Satyrus leechi Grum-Grshimailo, 1890

SUBS Karanasa leechi leechi (Grum-Grshimailo, 1890)
SUBS Karanasa leechi intermedius Grum-Grshimailo, 1890
SUBS Karanasa leechi erubescens Avinoff & Sweadner, 1951
SUBS Karanasa leechi alitchura Avinoff & Sweadner, 1951
SUBS Karanasa leechi centralis Avinoff & Sweadner, 1951
SUBS Karanasa leechi gregorii Avinoff & Sweadner, 1951
SUBS Karanasa leechi mihmana Avinoff & Sweadner, 1951
SUBS Karanasa leechi hunza Avinoff & Sweadner, 1951

SPEC Karanasa hoffmanni (Christoph, 1893)

ORIG Satyrus Huebneri var. Hoffmanni Christoph, 1893

SPEC Karanasa incerta Bogdanov, 1997
SPEC Karanasa straminea Bogdanov, 1997
SPEC Karanasa latifasciata (Grum-Grshimailo, 1902)

ORIG Satyrus regeli var. latifascia Grum-Grshimailo, 1902

SUBS Karanasa latifasciata latifasciata (Grum-Grshimailo, 1902)
SUBS Karanasa latifasciata obscurior Avinoff & Sweadner, 1951
SUBS Karanasa latifasciata occidentalis Avinoff & Sweadner, 1951

SPEC Karanasa regeli (Alphéraky, 1881)

ORIG Satyrus regeli Alphéraky, 1881

SUBS Karanasa regeli regeli (Alphéraky, 1881)
SUBS Karanasa regeli rueckbeili Avinoff & Sweadner, 1951
SUBS Karanasa regeli korlana Staudinger, 1901
SUBS Karanasa regeli aksuensis Avinoff & Sweadner, 1951

SPEC Karanasa eburnea Avinoff & Sweadner, 1951
SPEC Karanasa tancrei (Grum-Grshimailo, 1898)
SPEC Karanasa puengeleri (A. Bang-Haas, 1910)

ORIG Satyrus pungeleri A. Bang-Haas, 1910

SPEC Karanasa abramovi (Erschoff, 1884)

ORIG Satyrus abramovi Erschoff, 1884

SUBS Karanasa abramovi abramovi (Erschoff, 1884)
SUBS Karanasa abramovi regula (Staudinger, 1887)

ORIG Satyrus regeli var. regulus Staudinger, 1887
SSYN Eumenia regeli minutianus Fruhstorfer, 1911
SSYN Karanasa abramovi lactaea Avinoff & Sweadner, 1951

SUBS Karanasa abramovi naryna Avinoff & Sweadner, 1951
SUBS Karanasa abramovi striata Bogdanov, 2001

SPEC Karanasa kasakstana (O. Bang-Haas, 1936)

ORIG Satyrus regeli kasakstana O. Bang-Haas, 1936

SPEC Karanasa pamira (Staudinger, 1887)

ORIG Satyrus pamirus Staudinger, 1887

SUBS Karanasa pamira pamira (Staudinger, 1887)
SUBS Karanasa pamira ornata Avinoff & Sweadner, 1951
SUBS Karanasa pamira hohlbecki Avinoff & Sweadner, 1951
SUBS Karanasa pamira kotandari Wyatt, 1961
SUBS Karanasa pamira disjuncta Bogdanov, 2001

SPEC Karanasa haslundi Avinoff & Sweadner, 1951
SPEC Karanasa alpherakyi (Avinoff, 1910)

ORIG Satyrus alpherakyi Avinoff, 1910

SUBS Karanasa alpherakyi alpherakyi (Avinoff, 1910)
SUBS Karanasa alpherakyi kafir Avinoff & Sweadner, 1951
SUBS Karanasa alpherakyi twomeyi Wyatt, 1961
SUBS Karanasa alpherakyi titan Wyatt & Omoto, 1966
SUBS Karanasa alpherakyi transitoria Bogdanov, 2001

SPEC Karanasa moorei (Evans, 1912)

SUBS Karanasa moorei moorei (Evans, 1912)
SUBS Karanasa moorei haarlovi Avinoff & Sweadner, 1951
SUBS Karanasa moorei gilgitica Tytler, 1926
SUBS Karanasa moorei dubia Avinoff & Sweadner, 1951

SPEC Karanasa modesta Moore, 1893

SUBS Karanasa modesta modesta Moore, 1893
SUBS Karanasa modesta gemina Avinoff & Sweadner, 1951
SUBS Karanasa modesta baltorensis Avinoff & Sweadner, 1951

SPEC Karanasa astorica Tytler, 1926

SUBS Karanasa astorica astorica Tytler, 1926
SUBS Karanasa astorica expressa Avinoff & Sweadner, 1951
SUBS Karanasa astorica balti Tytler, 1926

SSYN Karanasa leechi Moore, 1893

SUBS Karanasa astorica pallida Tytler, 1926

SPEC Karanasa huebneri (Felder & Felder, 1867)

ORIG Satyrus huebneri Felder & Felder, 1867

SPEC Karanasa pupilata Tytler, 1926
SPEC Karanasa safeda Tytler, 1926
SPEC Karanasa cadesia (Moore, 1875)

ORIG Hipparchia cadesia Moore, 1875

SPEC Karanasa rohtanga Avinoff & Sweadner, 1951
SPEC Karanasa goergneri Eckweiler, 1990

GENU Neominois Scudder, 1875

SPEC Neominois ridingsii (W. H. Edwards, 1865)

SUBS Neominois ridingsii minimus (Austin, 1986)
SUBS Neominois ridingsii wyomingo (Scott, 1998)
SUBS Neominois ridingsii ridingsii (W. H. Edwards, 1865)
SUBS Neominois ridingsii curicata (M. Fisher, Scott & Garhart, 2008)
SUBS Neominois ridingsii coloalbiterra (Garhart & M. Fisher, 2008)
SUBS Neominois ridingsii neomexicanus (Austin, 1986)
SUBS Neominois ridingsii stretchii (W. H. Edwards, 1870)
SUBS Neominois ridingsii pallidus (Austin, 1986)

SPEC Neominois carmen A. Warren, Austin, Llorente, Luis & Vargas, 2008

GENU Oeneis Hübner, 1819

GSYN Chionobas Boisduval, 1832

SPEC Oeneis melissa (Fabricius, 1775)

ORIG Papilio melissa Fabricius, 1775

SUBS Oeneis melissa melissa (Fabricius, 1775)
SUBS Oeneis melissa karae Kusnezov, 1925
SUBS Oeneis melissa gibsoni Holland, 1931
SUBS Oeneis melissa orientalis Kurentzov, 1970
SUBS Oeneis melissa daizetsuzana Matsumura, 1926
SUBS Oeneis melissa semidea (Say, 1828)
SUBS Oeneis melissa semplei Holland, 1931
SUBS Oeneis melissa assimilis Butler, 1868
SUBS Oeneis melissa gibsoni Holland, 1931
SUBS Oeneis melissa beanii Elwes, 1893
SUBS Oeneis melissa lucilla Barnes & McDunnough, 1918

SPEC Oeneis tunga Staudinger, 1894
SPEC Oeneis jutta (Hübner, 1806)

SUBS Oeneis jutta jutta (Hübner, 1806)
SUBS Oeneis jutta alaskensis Holland, 1900
SUBS Oeneis jutta leussleri Bryant, 1935
SUBS Oeneis jutta harperi dos Passos, 1976
SUBS Oeneis jutta chermocki Wyatt, 1965
SUBS Oeneis jutta reducta McDunnough, 1929
SUBS Oeneis jutta ridingiana F. & R. Chermock, 1940
SUBS Oeneis jutta ascerta Masters & Sorenson, 1968
SUBS Oeneis jutta terranovae dos Passos, 1935
SUBS Oeneis jutta akoene Belik & Yakovlev, 1998
SUBS Oeneis jutta sibirica Kurentzov, 1970
SUBS Oeneis jutta balderi (Geyer, 1837)

SPEC Oeneis fulla (Eversmann, 1851)
SPEC Oeneis magna Graeser, 1888

SUBS Oeneis magna magna Graeser, 1888
SUBS Oeneis magna dubia Elwes, 1899
SUBS Oeneis magna magadanica Kurentzov, 1970
SUBS Oeneis magna kamtschatica Kurentzov, 1970
SUBS Oeneis magna pupavkini Korshunov, 1995
SUBS Oeneis magna kurentzovi Murayama, 1973
SUBS Oeneis magna uchangi Im, 1988

SPEC Oeneis glacialis (Moll, 1785)

ORIG Papilio glacialis Moll, 1785
SSYN Papilio aello Hübner, 1803-1804

SPEC Oeneis norna (Thunberg, 1791)

ORIG Papilio norna Thunberg, 1791

SUBS Oeneis norna norna (Thunberg, 1791)
SUBS Oeneis norna altaica Elwes, 1899
SUBS Oeneis norna chione Austaut, 1911
SUBS Oeneis norna aretuhusoides Lukhtanov, 1989
SUBS Oeneis norna rishirizana Matsumura, 1929
SUBS Oeneis norna takanonis Matsumura, 1909
SUBS Oeneis norna tundra Bang-Haas, 1912
SUBS Oeneis norna radnaevi Churkin, 1999
SUBS Oeneis norna rosovi Kurentzov, 1970
SUBS Oeneis norna tshukota Korshunov, 1998
SUBS Oeneis norna asamana Matsumura, 1919
SUBS Oeneis norna sugitanii Shirôzu, 1952
SUBS Oeneis norna hilda (Quensel, 1791)

SPEC Oeneis polixenes (Fabricius, 1775)

ORIG Papilio polixenes Fabricius, 1775

SUBS Oeneis polixenes polixenes (Fabricius, 1775)
SUBS Oeneis polixenes beringiana Kurentzov, 1970
SUBS Oeneis polixenes peartiae (Edwards, 1897)
SUBS Oeneis polixenes katahdin (Newcomb, 1901)
SUBS Oeneis polixenes subhyalina (Curtis, 1835)
SUBS Oeneis polixenes yukonensis Gibson, 1920
SUBS Oeneis polixenes brucei (Edwards, 1891)
SUBS Oeneis polixenes woodi Troubridge & Parshall, 1988
SUBS Oeneis polixenes paior Lukhtanov, 1989
SUBS Oeneis polixenes antonovae Lukhtanov, 1989

SPEC Oeneis actaeoides Lukhtanov, 1989

SUBS Oeneis actaeoides actaeoides Lukhtanov, 1989
SUBS Oeneis actaeoides czekanowskii Lukhtanov, 1989

SPEC Oeneis philipi Troubridge, 1988
SPEC Oeneis alpina Kurentzov, 1970

SUBS Oeneis alpina alpina Kurentzov, 1970
SUBS Oeneis alpina excubitor Troubridge, Philip, Scott & Schepard, 1982

SPEC Oeneis elwesi Staudinger, 1901

SUBS Oeneis elwesi elwesi Staudinger, 1901
SUBS Oeneis elwesi tannuola O. Bang-Haas, 1927
SUBS Oeneis elwesi ulugchemi Korshunov, 1995

SPEC Oeneis mulla Staudinger, 1881
SPEC Oeneis ammosovi Dubatov & Korshunov, 1988
SPEC Oeneis hora Grum-Grshimailo, 1888
SPEC Oeneis aktashi Lukhtanov, 1984
SPEC Oeneis ammon Elwes, 1899
SPEC Oeneis bore (Schneider, 1792)

SUBS Oeneis bore bore (Schneider, 1792)
SUBS Oeneis bore pansa Christoff, 1893
SUBS Oeneis bore arasaguna Austaut, 1911
SUBS Oeneis bore patrushevae Korshunov, 1985
SUBS Oeneis bore mackinleyensis dos Passos, 1949
SUBS Oeneis bore hanburyi Watkins, 1928
SUBS Oeneis bore taygete Geyer, 1830
SUBS Oeneis bore gaspeensis dos Passos, 1949
SUBS Oeneis bore fordi dos Passos, 1949
SUBS Oeneis bore edwardsi dos Passos, 1949

SPEC Oeneis nevadensis (Felder & Felder, 1866)

SUBS Oeneis nevadensis nevadensis (Felder & Felder, 1866)
SUBS Oeneis nevadensis iduna (Edwards, 1874)
SUBS Oeneis nevadensis gigas Butler, 1868

SPEC Oeneis macounii (Edwards, 1885)
SPEC Oeneis tanana A. Warren & Nakahara, 2016
SPEC Oeneis chryxus (Doubleday & Hewitson, 1849)

SUBS Oeneis chryxus chryxus (Doubleday & Hewitson, 1849)
SUBS Oeneis chryxus stanislaus Hovanitz, 1937
SUBS Oeneis chryxus valerata Burdick, 1958
SUBS Oeneis chryxus strigulosa McDunnough, 1934
SUBS Oeneis chryxus calais (Scudder, 1865)
SUBS Oeneis chryxus caryi Dyar, 1904
SUBS Oeneis chryxus altacordillera Scott, 2006 SUBS Oeneis chryxus socorro R. Holland, 2010 SUBS Oeneis chryxus ivallda (Mead, 1878)

SPEC Oeneis alberta Elwes, 1893

SUBS Oeneis alberta alberta Elwes, 1893
SUBS Oeneis alberta oslari Skinner, 1911
SUBS Oeneis alberta capulinensis F.M. Brown, 1970
SUBS Oeneis alberta daura (Strecker, 1894)
SUBS Oeneis alberta ojibwe Scott & Kondla, 2006

SPEC Oeneis buddha Grum-Grshimailo, 1891

SUBS Oeneis buddha buddha Grum-Grshimailo, 1891
SUBS Oeneis buddha dejeani O. Bang-Haas, 1939
SUBS Oeneis buddha grieshuberi Lukhtanov & Eitschberger, 2000
SUBS Oeneis buddha kincli Kocman, 1994
SUBS Oeneis buddha frankenbachi Lukhtanov & Eitschberger, 2000
SUBS Oeneis buddha pygmea Gross, 1970
SUBS Oeneis buddha brahma O. Bang-Haas, 1913
SUBS Oeneis buddha garhwalica Tytler, 1926

SPEC Oeneis oeno (Boisduval, 1832)

GENU Davidina Oberthür, 1879

GSYN Leechia Röber, [1907]
GSYN Sinosatyrus Lee
GSYN Protoeneis Gorbunov, 2001

SPEC Davidina armandi Oberthür, 1879

SSYN Leechia alticola Röber, [1907]
SSYN Sinosatyrus beyingenis Lee, 1988

SPEC Davidina diluta (Lukhtanov, 1994)
SPEC Davidina lederi (Alpheraky, 1897)

SSYN Oeneis sapozhnikovi Korshunov, 1982

SPEC Davidina mongolica (Oberthür, 1876)

SUBS Davidina mongolica mongolica (Oberthür, 1876)

SSYN Oeneis mongolica tsingtaua Austaut, 1911
SSYN Oeneis mongolica mandschurica O. Bang-Haas, 1939

SUBS Davidina mongolica hoenei (Gross, 1970)
SUBS Davidina mongolica coreana (Matsumura, 1927)
SUBS Davidina mongolica walkyria (Fixsen, 1887)
SUBS Davidina mongolica hallasanensis (Murayama, 1991)

SPEC Davidina nanna (Ménétriés, 1859)

SUBS Davidina nanna nanna (Ménétriés, 1859)

SSYN Oeneis hulda Staudinger, 1887
SSYN Oeneis nanna coriacea Seitz, 1909

SUBS Davidina nanna anna (Austaut, 1911)

SSYN Oeneis brunhilda A. Bang-Haas, 1912
SSYN Oeneis anna burjatica Korshunov et Nikolaev, 2002

SUBS Davidina nanna dzhugdzhuri (Shjeljuzhko, 1929)
SUBS Davidina nanna dzhulukuli (Korshunov, 1998)
SUBS Davidina nanna jakutski (Korshunov, 1998)
SUBS Davidina nanna taimyrica (Lukhtanov & Eitschberger, 2001)

SPEC Davidina sculda (Eversmann, 1851)

SUBS Davidina sculda sculda (Eversmann, 1851)

SSYN Oeneis velleda Austaut, 1911

SUBS Davidina sculda pseudosculda (Korshunov, 1977)
SUBS Davidina sculda pumila (Staudinger, 1892)
SUBS Davidina sculda vadimi (Korshunov, 1995)

SPEC Davidina tarpeia (Pallas, 1771)

SUBS Davidina tarpeia tarpeia (Pallas, 1771)

SSYN Papilio celimene Cramer, 1782
SSYN Oeneis vacuna Grum-Grshimailo, [1891]
SSYN Oeneis tarpeia rozhdestvenskyi Korb & Yakovlev, 1997
SSYN Oeneis tarpeia ukokana Korb, 1998
SSYN Oeneis tarpeia baueri Lukhtanov & Eitschberger, 2000

SUBS Davidina tarpeia grossi (Eitschberger & Lukhtanov, 1994)

SPEC Davidina uhleri (Reakirt, 1866)

SUBS Davidina uhleri uhleri (Reakirt, 1866)

SSYN Chionobas uhleri obscura (W. H. Edwards, 1892

SUBS Davidina uhleri reinthali (F.M. Brown, 1953)
SUBS Davidina uhleri varuna (Edwards, 1882)
SUBS Davidina uhleri nahanni (Dyar, 1904)
SUBS Davidina uhleri cairnesi (Gibson, 1920)

SSYN Oeneis nanna kluanensis Hassler, 2002

SPEC Davidina urda (Eversmann, 1847)

SUBS Davidina urda urda (Eversmann, 1847)

SSYN Oeneis urda umbra Staudinger, 1892
SSYN Oeneis urda laeta Austaut, 1908
SSYN Oeneis urda trybomi Bryk, 1946

SUBS Davidina urda tschiliensis (O. Bang-Haas, 1933)
SUBS Davidina urda monteviri (Bryk, 1946)

GENU Paroeneis Moore, 1893

SPEC Paroeneis pumilus (Felder & Felder, 1867)

ORIG Chionobas pumilus Felder & Felder, 1867

SPEC Paroeneis sikkimensis (Staudinger, 1889)
SPEC Paroeneis palearcticus (Staudinger, 1889)

SUBS Paroeneis palearcticus palearcticus (Staudinger, 1889)
SUBS Paroeneis palearcticus nanshanicus (Grum-Grshimailo, 1902)
SUBS Paroeneis palearcticus buddha Bang-Haas, 1927
SUBS Paroeneis palearcticus iole (Leech, 1892)
SUBS Paroeneis palearcticus atuntsensis (Gross, 1958)
SUBS Paroeneis palearcticus auloceroides Huang, 1999

SPEC Paroeneis parapumilus Huang, 2001
SPEC Paroeneis bicolor (Seitz, 1909)
SPEC Paroeneis grandis Riley, 1923

GENU Arethusana de Lesse, 1951

SPEC Arethusana arethusa (Schiffermüller, 1775)

ORIG Papilio arethusa Schiffermüller, 1775

SUBS Arethusana arethusa arethusa (Schiffermüller, 1775)
SUBS Arethusana arethusa heptapotamica (Stauder, 1924)
SUBS Arethusana arethusa pontica (Heyne, 1895)
SUBS Arethusana arethusa hakkarica Koçak, 1975

SPEC Arethusana aksouali Wyatt, 1952

GENU Brintesia Fruhstorfer, 1911

GSYN Oreas Hübner, 1806 (reject.)
GSYN Oreas Oken, 1815 (reject.)

SPEC Brintesia circe (Fabricius, 1775)

ORIG Papilio circe Fabricius, 1775
SSYN Papilio proserpina Denis & Schiffermüller, 1775

SUBS Brintesia circe circe (Fabricius, 1775)
SUBS Brintesia circe venusta (Fruhstorfer, 1909)
SUBS Brintesia circe asperomontana (Stauder, 1921)
SUBS Brintesia circe pannonia (Fruhstorfer, 1917)

GENU Minois Hübner, 1819

SPEC Minois dryas (Scopoli, 1763)

ORIG Papilio dryas Scopoli, 1763
SSYN Papilio phaedra Linnaeus, 1764

SUBS Minois dryas dryas (Scopoli, 1763)
SUBS Minois dryas septentrionalis Wnukowsky, 1929
SUBS Minois dryas bipunctatus (Motschulsky, 1861)
SUBS Minois dryas shaanxiensis Qian

SPEC Minois nagasawae (Matsumura, 1906)
SPEC Minois paupera (Alphéraky, 1888)
SPEC Minois aurata (Oberthür, 1909)

GENU Chazara Moore, 1893

GSYN Philareta Moore, 1893

SPEC Chazara briseis (Linnaeus, 1764)

ORIG Papilio briseis Linnaeus, 1764

SUBS Chazara briseis briseis (Linnaeus, 1764)
SUBS Chazara briseis major (Oberthür, 1876)

SSYN Satyrus briseis cretus Verity, 1913

SUBS Chazara briseis meridionalis (Staudinger, 1886)
SUBS Chazara briseis magna Heyne, 1894
SUBS Chazara briseis armena Jachontov, 1911
SUBS Chazara briseis hyrcana (Staudinger, 1886)
SUBS Chazara briseis fergana (Staudinger, 1886)
SUBS Chazara briseis saga (Fruhstorfer, 1909)
SUBS Chazara briseis lyrnessus (Fruhstorfer, 1908)

SPEC Chazara heydenreichi (Kindermann, 1853)

SUBS Chazara heydenreichi heydenreichi (Kindermann, 1853)
SUBS Chazara heydenreichi hegesander Fruhstorfer, 1910
SUBS Chazara heydenreichi nana Rühl, 1895

SPEC Chazara persephone (Hübner, 1805)

ORIG Eumenis persephone Hübner, 1805
SSYN Papilio anthe Ochsenheimer, 1807
SSYN Satyrus hanifa Herrich-Schaeffer, 1850

SUBS Chazara persephone persephone (Hübner, 1805)
SUBS Chazara persephone transies Zerny, 1932
SUBS Chazara persephone pseudoenervata Lukhtanov, 1999

SPEC Chazara enervata (Alpheraky, 1882)
SPEC Chazara kaufmanni (Erschoff, 1874)

SUBS Chazara kaufmanni kaufmanni (Erschoff, 1874)
SUBS Chazara kaufmanni sieversi (Christoph, 1885)
SUBS Chazara kaufmanni obscurior (Staudinger, 1887)
SUBS Chazara kaufmanni sartha (Staudinger, 1886)

SPEC Chazara eitschbergeri Lukhtanov, 1999
SPEC Chazara staudingeri (Bang-Haas, 1882)

SUBS Chazara staudingeri staudingeri (Bang-Haas, 1882)
SUBS Chazara staudingeri tadjika (Grum-Grshimailo, 1890)
SUBS Chazara staudingeri gultschensis (Grum-Grshimailo, 1888)

SPEC Chazara rangontavica Shchetkin, 1981
SPEC Chazara bischoffii (Herrich-Schaeffer, 1846)

ORIG Satyrus bischoffi Herrich-Schaeffer, 1846

SPEC Chazara prieuri (Pierret, 1837)

ORIG Satyrus prieuri Pierret, 1837
SSYN Satyrus prieuri kebira Wyatt, 1952

GENU Pseudochazara de Lesse, 1951

SPEC Pseudochazara hippolyte (Esper, 1784)

SUBS Pseudochazara hippolyte hippolyte (Esper, 1784)
SUBS Pseudochazara hippolyte mercurius Staudinger, 1887
SUBS Pseudochazara hippolyte dorriesi (Bang-Haas, 1933)

SPEC Pseudochazara williamsi Romei, 1927
SPEC Pseudochazara pallida (Staudinger, 1901)
SPEC Pseudochazara euxina (Kusnetsov, 1909)
SPEC Pseudochazara turkestana (Grum.Grshimailo, 1893)

SUBS Pseudochazara turkestana turkestana (Grum.Grshimailo, 1893)
SUBS Pseudochazara turkestana tarbagata (Staudinger, 1901)
SUBS Pseudochazara turkestana sagina (Rühl, 1894)
SUBS Pseudochazara turkestana esquilinus (Fruhstorfer, 1911)

SPEC Pseudochazara panjshira Wyatt & Omoto, 1966

SUBS Pseudochazara panjshira panjshira Wyatt & Omoto, 1966
SUBS Pseudochazara panjshira kopetdaghi Dubatolov, 1989
SUBS Pseudochazara panjshira badachshana Wyatt & Omoto, 1966

SPEC Pseudochazara pakistana Gross, 1978
SPEC Pseudochazara pelopea (Klug, 1832)

ORIG Hipparchia pelopea Klug, 1832

SUBS Pseudochazara pelopea pelopea (Klug, 1832)
SUBS Pseudochazara pelopea persica (Christoph, 1877)
SUBS Pseudochazara pelopea tekkensis (Heyne, 1895)

SPEC Pseudochazara schahrudensis (Staudinger, 1881)
SPEC Pseudochazara nukatli Bogdanov, 2000
SPEC Pseudochazara mamurra (Herrich-Schäffer, 1844)

ORIG Satyrus mamurra Herrich-Schäffer, 1844

SUBS Pseudochazara mamurra mamurra (Herrich-Schäffer, 1844)
SUBS Pseudochazara mamurra birgit Gross, 1978

SPEC Pseudochazara alpina (Staudinger, 1878)
SPEC Pseudochazara guriensis (Staudinger, 1878)
SPEC Pseudochazara beroe (Herrich-Schäffer, 1844)

SUBS Pseudochazara beroe beroe (Herrich-Schäffer, 1844)
SUBS Pseudochazara beroe rhena (Herrich-Schäffer, 1852)
SUBS Pseudochazara beroe aurantiaca (Staudinger, 1871)

SPEC Pseudochazara schakuhensis (Staudinger, 1881)
SPEC Pseudochazara mniszechii (Herrich-Schäffer, 1851)

ORIG Eumenis mniszechii Herrich-Schäffer, 1851

SUBS Pseudochazara mniszechii mniszechii (Herrich-Schäffer, 1851)
SUBS Pseudochazara mniszechii caucasica (Lederer, 1864)
SUBS Pseudochazara mniszechii watsoni Clench & Shoumatoff, 1956

SPEC Pseudochazara geyeri (Herrich-Schäffer, 1846)

ORIG Satyrus geyeri Herrich-Schäffer, 1846

SUBS Pseudochazara geyeri geyeri (Herrich-Schäffer, 1846)
SUBS Pseudochazara geyeri karsicola Gross, 1978
SUBS Pseudochazara geyeri occidentalis (Rebel & Zerny, 1931)

SPEC Pseudochazara daghestana Holik, 1955

SUBS Pseudochazara daghestana daghestana Holik, 1955
SUBS Pseudochazara daghestana savalanica Gross & Ebert, 1975

SPEC Pseudochazara kanishka Aussem, 1980
SPEC Pseudochazara thelephassa (Geyer, 1827)

ORIG Eumenis thelephassa Geyer, 1827

SPEC Pseudochazara anthelea (Hübner, 1824)

SUBS Pseudochazara anthelea anthelea (Hübner, 1824)
SUBS Pseudochazara anthelea schawerdae (Fruhstorfer, 1908)
SUBS Pseudochazara anthelea selcuki Van Oorschot, 1987

SPEC Pseudochazara amalthea (Frivaldsky, 1845)
SPEC Pseudochazara atlantis (Austaut, 1905)

ORIG Satyrus atlantis Austaut, 1905
SSYN Satyrus mniszechi var. maroccana Meade-Waldo, 1906
SSYN Satyrus atlantis colini Wyatt, 1952
SSYN Pseudochazara atlantis benderi Weiss, 1979

SPEC Pseudochazara baldiva (Moore, 1865)
SPEC Pseudochazara droshica (Tytler, 1926)

SUBS Pseudochazara droshica droshica (Tytler, 1926)
SUBS Pseudochazara droshica rajevskyi Tshikolovets, 1996

SPEC Pseudochazara lehana (Moore, 1878)

SUBS Pseudochazara lehana lehana (Moore, 1878)
SUBS Pseudochazara lehana riegeri Tshikolovets, 2005

SPEC Pseudochazara graeca (Staudinger, 1870)
SPEC Pseudochazara lydia (Staudinger, 1878)

SUBS Pseudochazara lydia lydia (Staudinger, 1878)
SUBS Pseudochazara lydia obscura (Staudinger, 1878)
SUBS Pseudochazara lydia aurora Eckweiler & Rose, 1989

SPEC Pseudochazara cingovskii Gross, 1973
SPEC Pseudochazara orestes de Prins & Poorten, 1981
SPEC Pseudochazara amymone Brown, 1976
SPEC Pseudochazara porphyritica Clench & Shoumatoff, 1956

GENU Satyrus Latreille, 1810

SPEC Satyrus actaea (Esper, 1780)

ORIG Papilio actaea Esper, 1780

SPEC Satyrus ferula (Fabricius, 1793)

ORIG Papilio ferula Fabricius, 1793

SUBS Satyrus ferula ferula (Fabricius, 1793)
SUBS Satyrus ferula atlantea (Verity, 1927)

ORIG Nytha actaea atlantea Verity, 1927 (repl. maroccana Lucas)
SSYN Satyrus actaea f. maroccana D. Lucas, 1920
SSYN Satyrus actaea f. meknesensis Strand, 1927
SSYN Satyrus actaea simillima Rothschild, 1931

SUBS Satyrus ferula altaica Grum-Grshimailo, 1893
SUBS Satyrus ferula liupiuschani O. Bang-Haas, 1933
SUBS Satyrus ferula medvedevi Korshunov, 1996
SUBS Satyrus ferula cordulina Staudinger, 1886
SUBS Satyrus ferula alaica Staudinger, 1886
SUBS Satyrus ferula rickmersi van Rosen, 1921
SUBS Satyrus ferula penketia Fruhstorfer, 1908
SUBS Satyrus ferula serva Fruhstorfer, 1909

SPEC Satyrus stheno Grum-Grshimailo, 1887
SPEC Satyrus virbius Herrich-Schäffer, 1843
SPEC Satyrus amasinus Staudinger, 1861
SPEC Satyrus iranicus Schwingenschuss, 1939

SUBS Satyrus iranicus iranicus Schwingenschuss, 1939
SUBS Satyrus iranicus kyros Gross & Ebert, 1975

SPEC Satyrus effendi Nekrutenko, 1989
SPEC Satyrus daubi Gross & Ebert, 1975
SPEC Satyrus nana Staudinger, 1886
SPEC Satyrus pimpla Felder & Felder, 1867

SUBS Satyrus pimpla pimpla Felder & Felder, 1867
SUBS Satyrus pimpla tajik Clench & Shoumatoff, 1956

SPEC Satyrus orphei Shchetkin, 1985
SPEC Satyrus favonius Staudinger, 1891
SPEC Satyrus parthicus Lederer, 1869

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